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Articles by Juha-Pekka Hotanen

Category: Research article

article id 1005, category Research article
Marjut Turtiainen, Jari Miina, Kauko Salo, Juha-Pekka Hotanen. (2013). Empirical prediction models for the coverage and yields of cowberry in Finland. Silva Fennica vol. 47 no. 3 article id 1005. https://doi.org/10.14214/sf.1005
Highlights: The site fertility significantly affected the abundance of cowberry on mineral soils, spruce mires and pine mires; The stand basal area and dominant tree species were among the most important forest structural predictors in the model for the coverage; In the cowberry yield model developed for mineral soil sites, the stand basal area and coverage of cowberry plants were statistically significant predictors.
Empirical models for the coverage and berry yield of cowberry (Vaccinium vitis-idaea L.) were developed using generalized linear mixed models (GLMMs). The percentage coverage of cowberry was predicted as a function of site and stand characteristics using data from the Finnish National Forest Inventory (NFI) in 1995. The average annual yield, including the between-year variation in the yield, was predicted as a function of percentage coverage and stand characteristics using permanent experimental plots (MASI) established in different areas of Finland and measured in 2001-2012. The model for cowberry yields (Model 2) was developed for mineral soil forests. The model for the coverage (Model 1) was constructed so that it considers both mineral soil sites and also many other sites where cowberry occurs in the field layer. According to Model 1, the site fertility significantly affected the abundance of cowberry on mineral soils, spruce mires and pine mires. The stand basal area and dominant tree species were among the most important forest structural predictors in Model 1. The site fertility was not a significant predictor in the cowberry yield model. Instead, the stand basal area and coverage of cowberry plants were found to be statistically significant predictors in Model 2. The estimated models were used to predict the cowberry coverage, average annual yield and its 95 % confidence interval along with stand development. The models of this study can be used for multi-objective forest planning purposes.
  • Turtiainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: marjut.turtiainen@uef.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: jari.miina@metla.fi
  • Salo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: kauko.salo@metla.fi
  • Hotanen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: juha-pekka.hotanen@metla.fi
article id 181, category Research article
Jari Miina, Juha-Pekka Hotanen, Kauko Salo. (2009). Modelling the abundance and temporal variation in the production of bilberry (Vaccinium myrtillus L.) in Finnish mineral soil forests. Silva Fennica vol. 43 no. 4 article id 181. https://doi.org/10.14214/sf.181
Empirical models for the abundance and berry yield of V. myrtillus were constructed using generalized linear mixed model (GLMM) techniques. The percentage coverage of bilberry was predicted as a function of site and stand characteristics using the permanent sample plots of the National Forest Inventory (NFI) in 1995. The number of bilberries was predicted as a function of percentage coverage and stand characteristics using a sub-sample of the NFI plots in North Karelia. The between-year variation in the bilberry yield was analysed using the permanent experimental plots (MASI) established in different areas of Finland and measured in 2001–2007. The highest coverage of bilberry was found on mesic heath sites; on sub-xeric and herb-rich heath sites the values were 62% of that for mesic sites. The decreasing effect of deciduous trees (compared to spruce) was significant only on herb-rich heath sites. The coverage increased along with stand development up to certain stand ages and basal areas. The bilberry yields were higher in pine-dominated stands than in spruce-dominated ones. In spruce stands, the coverage of bilberry and stand basal area significantly affected the number of berries, whereas in pine stands only the coverage was a significant predictor. In the MASI data, the bilberry yield of pine stands was two times higher than that of spruce stands; however, the between-year variation in bilberry yield was higher in the spruce than in the pine stands. The estimated models were used to predict the bilberry coverage and yield along with stand development. On mesic heath sites in southern Finland (1200 dd.), the predicted annual yield of bilberry was about 25 kg ha–1 (95% confidence interval 9–73 kg ha–1) in a mature pine stand and about 10 kg ha–1 (3–35 kg ha–1) in a mature spruce stand. The models can be included in stand simulators, where they would facilitate the prediction of bilberry abundance and yields for silvicultural and forest planning purposes.
  • Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: jari.miina@metla.fi (email)
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Salo, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 352, category Research article
Juha-Pekka Hotanen, Matti Maltamo, Antti Reinikainen. (2006). Canopy stratification in peatland forests in Finland. Silva Fennica vol. 40 no. 1 article id 352. https://doi.org/10.14214/sf.352
Abundance and species number of the tree and shrub vegetation in different canopy layers were analysed according to site quality class and drainage succession phase on permanent sample plots on spruce mires (n = 268) and pine mires (n = 628) in the Finnish National Forest Inventory in 1995. The abundances based on the crown coverage were compared with the abundances based on the parallel basal area of the tree stand. The canopy coverages and species number for peatland forests were also compared with those for mineral soil forests on the permanent sample plots (n = 1725) in 1995. In general, effective temperature sum correlated positively, although not very strongly, with the coverages and species number in most of the canopy layers, as well as with the mean range of the diameter distribution. The effects of both site quality class and drainage phase were stronger on pine mires than on spruce mires, most probably due to the longer fertility gradient and large potential free growing space in the former group. On pine mires, drainage increased the abundances and species number in the different canopy layers, as well as the structural inequality of the tree stands. On spruce mires, the increase was principally allocated to the abundances of the dominant and intermediate tree layers. The correlations between the total crown coverage of the tree layers and stand basal area were r = 0.45 for spruce mires and r = 0.70 for pine mires. Compared to mineral soil forests, in addition to having a higher abundance of Betula pubescens, the dominant layer was not as pronounced in peatland forests. On spruce mires, the coverage of the shrub layer on mesotrophic and meso-oligotrophic sites was higher than that in mineral soil forests. The average species number in different canopy layers did not differ significantly between spruce mires and mineral soil forests in corresponding site quality classes. On pine mires, the species number was generally lower (except for the mesotrophic sites) than that in corresponding mineral soil forests.
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: juha-pekka.hotanen@metla.fi (email)
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Reinikainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
article id 512, category Research article
Juha-Pekka Hotanen. (2003). Multidimensional site description of peatlands drained for forestry. Silva Fennica vol. 37 no. 1 article id 512. https://doi.org/10.14214/sf.512
Sites (n = 119) on drained mires located in the southern aapa mire zone in Finland were analysed by multivariate techniques. The compositional trends of the understorey vegetation were analysed by means of hybrid multidimensional scaling (HMDS). In addition to field classification, two-way indicator species analysis (TWINSPAN) and flexible unweighted paired group arithmetic average (FUPGMA) classifications were used. The 1st HMDS dimension primarily reflected variation along a gradient from spruce mire influence to hummock-level bog influence. Variation in nutrient status was also connected to this gradient. Factors underlying the 2nd dimension were variation in nutrient status and drainage succession (moisture). Some sample plots representing herb-rich or Molinia-rich types were separated along the 3rd dimension. The variation in understorey vegetation (i.e. the ordination space) showed high maximum correlation with stand volume r = 0.81, mean annual stand volume increment r = 0.76, and post-drainage dominant height r = 0.75. The covariation between the vegetation and peat bulk density in both the 0–10 and 10–20 cm peat layers was also strong: r = 0.55 and r = 0.80. The correlations for Hv.Post were 0.64 and 0.81, respectively. Of the total macronutrient concentrations, phosphorus (r = 0.73, r = 0.75) and nitrogen (r = 0.59, r = 0.64) were the most strongly correlated with species composition. The environmental sample variables were also presented by the vegetation units of numerical classification. Most of the recorded variables, including nutrient amounts (kg ha–1), were examined in site quality (fertility) classes by succession phases as well. Border variants or transitional forms of the site types were common. Additional vegetation criteria (e.g. surface-water influence) more closely defined the ecology of the site. In addition to the site quality classes, a considerable amount of information about the tree stands, vegetation diversity and peat properties was associated with the separation of the succession phases, i.e. in this study transforming (phase II) vs. transformed (final phase III) sites. In conclusion, the actual vegetation appeared to well reflect various aspects of the ecological conditions, even in labile communities of commercial forests on drained peatlands.
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Centre, P.O.Box 68, FIN-80101 Joensuu, Finland ORCID ID:E-mail: juha-pekka.hotanen@metla.fi (email)

Category: Research note

article id 1573, category Research note
Marjut Turtiainen, Jari Miina, Kauko Salo, Juha-Pekka Hotanen. (2016). Modelling the coverage and annual variation in bilberry yield in Finland. Silva Fennica vol. 50 no. 4 article id 1573. https://doi.org/10.14214/sf.1573
Highlights: The highest bilberry coverage was found in mesic heath forests and fell forests; On peatlands the coverage was, on average, lower than on mineral soil sites; The approach introduced in this study to calculating annual berry yield indices is a promising way for estimating total annual bilberry yields over a given period of time.

The coverage of bilberry (Vaccinium myrtillus L.) was modelled as a function of site and stand characteristics using the permanent sample plots of the National Forest Inventory (NFI) (Model 1). The sample sites consisted of mineral soil forests as well as fells and peatland sites. Annual variation in the bilberry yield (Model 2) was analysed based on measurements over 2001–2014 in the permanent sample plots (so-called MASI plots) in various areas of Finland. We derived annual bilberry yield indices from the year effects of Model 2 and investigated whether these indices could be used to estimate annual variation in bilberry crops in Finland. The highest bilberry coverage was found in mesic heath forests and fell forests. On peatlands the coverage was, on average, lower than on mineral soil sites; the peatland sites with most bilberry coverage were meso-oligotrophic and oligotrophic spruce mires and oligotrophic pine mires. Our bilberry yield indices showed similar variation to those derived from the mean annual berry yields reported and calculated earlier using the MASI plots; the correlation between the indices was 0.795. This approach to calculating annual berry yield indices is a promising way for estimating total annual bilberry yields over a given period of time. Models 1 and 2 can be used in conjunction with the Miina et al.’s (2009) bilberry yield model when bilberry coverage, average annual yield and annual variation in the yield are to be predicted in forest planning.

  • Turtiainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: marjut.turtiainen@uef.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: jari.miina@luke.fi
  • Salo, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: kauko.salo@luke.fi
  • Hotanen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: juha-pekka.hotanen@luke.fi

Category: Article

article id 5572, category Article
Markku Lehtelä, Juha-Pekka Hotanen, Pentti Sepponen. (1996). Understorey vegetation in fresh and herb-rich upland forests in southwest Lapland. Silva Fennica vol. 30 no. 1 article id 5572. https://doi.org/10.14214/sf.a9217

Fresh and herb-rich upland forest sites in the north-western part of the central boreal vegetation zone in Finland were studied with respect to vegetation structure and vegetation-environment relationships (soil, stand characteristics). Two fresh heath vegetation data sets, one from the northern boreal zone and the other from the central boreal zone, were compared with the data of this study using multivariate methods.

The variation in heath forest vegetation within the climatically uniform area was mainly determined by the fertility of the soil (primarily Ca and Mg) and the stage of stand development. N, P and K content of the humus layer varied little between the vegetation classes. Fertile site types occurred, in general, on coarse-textured soils than infertile site types, may be due to the fact that the sample plots were located in various bedrock and glacial till areas, i.e. to sampling effects.

The place of the vegetational units of the study area in the Finnish forest site type system is discussed. The vegetation of the area has features in common with the northern boreal zone as well as the southern part of the central boreal vegetation zone. The results lend some support to the occurrence of a northern Myrtillus type or at least that intermediate form of fresh and herb-rich mineral soil sites commonly occur in the studied area. It is argued that the older name Dryopteris-Myrtillus type is more suitable than Geranium-Oxalis-Myrtillus type for herb-rich heath sites in the study area.

  • Lehtelä, ORCID ID:E-mail:
  • Hotanen, ORCID ID:E-mail:
  • Sepponen, ORCID ID:E-mail:

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