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Silva Fennica 1926-1997
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Acta Forestalia Fennica
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Articles containing the keyword 'growth rate'.

Category: Research article

article id 341, category Research article
Sanni Raiskila, Pekka Saranpää, Kurt Fagerstedt, Tapio Laakso, Mia Löija, Riitta Mahlberg, Leena Paajanen, Anne-Christine Ritschkoff. (2006). Growth rate and wood properties of Norway spruce cutting clones on different sites. Silva Fennica vol. 40 no. 2 article id 341. https://doi.org/10.14214/sf.341
The effect of growth rate on weight density and strength properties of three Norway spruce cutting clones growing on three different sites in different geographic locations was studied. The purpose was to follow variation in wood physical and mechanical properties and in quality between fast-growing clones grown in environments differing in nutritional and soil properties and climate within the boreal zone. The cloned trees had been selected on grounds of good growth, health and quality. The cuttings were collected from three-year-old seedlings and rooted. The rooted cuttings were planted in the 1970’s and they were on average 26 years old at a time of felling. The variation of weight density was studied within the annual ring and within the stem between the juvenile and mature wood from the pith to the bark with an X-ray densitometric method. The average annual ring width (and latewood proportion, %) varied between the clones from 2.92±1.36 mm (15.34%) to 3.30±1.25 mm (11.80%) and between the sites from 2.76±1.07 mm (14.71%) to 3.70±1.22 mm (13.29%). The mean weight density was 0.461±0.077 g cm–3 and latewood density 0.750±0.125 g cm–3 in this material. The mean modulus of elasticity was 9.88±1.43 GPa, modulus of rupture 67.51±11.50 MPa and weight density of the test samples (ρ12) 414±44 kg m–3 in mature wood. The parameters studied showed clearly that the environment had a large effect while the three clones differed from each other similarly in the different sites, e.g. the fastest growing clone was fastest on all sites.
  • Raiskila, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
  • Saranpää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: pekka.saranpaa@metla.fi (email)
  • Fagerstedt, Department of Biological and Environmental Sciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Laakso, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
  • Löija, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland ORCID ID:E-mail:
  • Mahlberg, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland ORCID ID:E-mail:
  • Paajanen, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland ORCID ID:E-mail:
  • Ritschkoff, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland ORCID ID:E-mail:
article id 439, category Research article
Christina Lundgren. (2004). Microfibril angle and density patterns of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 439. https://doi.org/10.14214/sf.439
Two Norway spruce nutrient optimisation trials, one in the north of Sweden and one in the south, were used to study the effects of intensive growth and fertilization on wood density and microfibril angle. Three different treatments and a control were available; daily irrigation, daily liquid fertilization and solid fertilization. The nutrient optimisation was based on foliage analysis and the solid fertilization essentially comprised the same amount of nutrients but was applied annually in solid form. Measurements of density and microfibril angle (MFA) were performed using X-ray diffraction. Growth rate, expressed as a transformation of annual ring width, was very important at the southern site when the effect of cambial maturation had been taken into account. Effects of both fertilization and irrigation remained strong and significant for density, and irrigation was a significant factor explaining MFA. At the northern site distance from pith was the dominant factor but the effect of growth rate was also strong and the treatment effect was significant for both density and MFA. The combination of higher MFA and decrease in density for fertilized trees resulted in a lower calculated strength of the wood. An over 100% increase in ring width only corresponded to approximately a 20% decrease in wood density and the production of wood dry matter was hence increased by treatments.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: christina.lundgren@spm.slu.se (email)
article id 600, category Research article
Qibin Yu, P. M. A. Tigerstedt, Matti Haapanen. (2001). Growth and phenology of hybrid aspen clones (Populus tremula L. x Populus tremuloides Michx.). Silva Fennica vol. 35 no. 1 article id 600. https://doi.org/10.14214/sf.600
Height, basal diameter, diameter at breast height, bud burst, and leaf development were recorded in a 5-year-old hybrid aspen clonal trial. The field trial consisted of four aspen hybrid clones (Populus tremula x Populus tremuloides) and one local P. tremula seedling source. Phenological traits were observed in the 3rd year. Growth patterns were recorded during the 3rd and 4th years. Phenological traits were explored in relation to hybrid vigor expressed as growth traits. Differences were observed for phenological and growth traits among hybrid clones and P. tremula. The growth period varied from 143–158 days for the four hybrid clones, and was 112 days for P. tremula. The correlation between growth period and yield was highly significant. The annual growth rate of height for the hybrids was 4.2 cm per 7 days (2.4 for P. tremula) in the 3rd year and 6.4 cm per 7 days (2.9 for P. tremula) in the 4th year. After 5 years, mean estimated stem volume of the hybrids was 3.9 times that of P. tremula. Significant clone by year interaction was observed for height, diameter, and volume growth. The hybrid vigor seems to be mainly attributable to a longer growth period.
  • Yu, Department of Plant Biology, P.O. Box 27, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail: qibin.yu@helsinki.fi (email)
  • Tigerstedt, Department of Plant Biology, P.O. Box 27, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Haapanen, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland ORCID ID:E-mail:
article id 666, category Research article
Rolf Pape. (1999). Influence of thinning on spiral grain in Norway spruce grown on highly productive sites in southern Sweden. Silva Fennica vol. 33 no. 1 article id 666. https://doi.org/10.14214/sf.666
Grain spirality was investigated in eight stands of Norway spruce (Picea abies (L.) Karst.) subjected to different thinning regimes. The dominating general pattern of spiral grain found in this study was typical for conifers, with a maximum of left-handed spirality close to the pith, which decreased towards the bark and sometimes changed to right-handed spiral grain in the outer growth rings. However, there was a large amount of between-tree variation in spiral grain. The effect of thinning on grain spirality was investigated by relating annual ring width to spiral grain, since thinning affects growth rate. A positive correlation between ring width and grain angle was found, but a considerable number of trees showed no or a negative correlation. A statistically significant effect of ring width was only found in five of the eight stands. Heavy thinnings, removing 60% of the basal area of a stand, considerably increased spiral grain, whereas the effects of light thinnings were inconsistent. These results support the findings of earlier studies indicating that spiral grain formation is under considerable genetic control, while its expression can be changed by silvicultural methods which affect growth rate.
  • Pape, Swedish University of Agricultural Sciences, Department of Forest Yield Research, P.O. Box 7061, S-750 07 Uppsala, Sweden ORCID ID:E-mail: rolf.pape@sprod.slu.se (email)

Category: Article

article id 5208, category Article
Matti Kärkkäinen. (1984). Effect of tree social status on basic density of Norway spruce. Silva Fennica vol. 18 no. 2 article id 5208. https://doi.org/10.14214/sf.a15387

The effect of growth rate on wood basic density in even-age Norway spruce (Picea abies (L.) H. Karst.) plantations was studied on the basis of samples collected from 53 stands; 30 trees were sampled in each stand. The prediction of basic density with the help of growth rate and some other tree characteristics could be improved if the social status of the tree was taken into account. Within a stand, the smaller trees had a lower density, while taller trees had a higher density than they should have had on the basis of growth rate alone.

The PDF includes a summary in Finnish.

  • Kärkkäinen, ORCID ID:E-mail:

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