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Articles containing the keyword 'tracheid'.

Category: Research article

article id 266, category Research article
Mikko Havimo, Juha Rikala, Jari Sirviö, Marketta Sipi. (2008). Distributions of tracheid cross-sectional dimensions in different parts of Norway spruce stems. Silva Fennica vol. 42 no. 1 article id 266. https://doi.org/10.14214/sf.266
Distributions of three cross-sectional dimensions: radial and tangential tracheid width, and cell wall thickness in different timber assortments of Norway spruce were investigated. Wood samples from a mature stand were measured with SilviScan. In the analysis, virtual trees were constructed from measurement data, and divided into three assortments: whole stem, top pulpwood and sawmill chips. Average values and distributions of the properties were calculated for all assortments, and distributions divided into earlywood and latewood across the whole tree assortment. There was considerable variation within latewood in all three cross-sectional dimensions, but variation in earlywood was slight in radial width and cell wall thickness. In earlywood, tangential tracheid width showed considerable internal variation, and the difference between earlywood and latewood in tangential width was small. Within-assortment variation of all three properties was larger than between assortments. We may conclude that only a moderate difference in pulp properties can be achieved by sorting raw material into sawmill chips and top pulpwood. Pulp fractionation into earlywood and latewood seems to be a more efficient method, since it gives classes with small within-class variation and distinct average properties. However, it should be kept in mind that the results are valid only in mature stands, where growth rate variation and juvenile wood content are small.
  • Havimo, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: mikko.havimo@helsinki.fi (email)
  • Rikala, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Sirviö, KCL, P.O. Box 70, FI-02151 Espoo, Finland ORCID ID:E-mail:
  • Sipi, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
article id 438, category Research article
Christina Lundgren. (2004). Cell wall thickness and tangential and radial cell diameter of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 438. https://doi.org/10.14214/sf.438
Two Norway spruce nutrient trials were used to evaluate the effects of fertilization and irrigation on transverse tracheid dimensions. Three different treatments and a control (C) were used; daily irrigation (I), daily liquid fertilization (IL) and an annual solid fertilization (F). The nutrient optimisation was based on foliage analysis and both liquid and solid fertilization essentially comprised the same amount of nutrients but the latter was applied annually in solid form. The cell measurements; cell wall thickness, radial and tangential cell widths, were obtained using image analysis (SilviScan at CSIRO, Melbourne, Australia). Mean annual cell wall thickness was decreased by fertilization (F and IL) on both sites whereas no effect of the irrigation on wall thickness could be detected. Radial cell width was increased by treatment at Flakaliden but at Asa the effect of irrigation and fertilization was reversed when the data structure i.e. development from pith and out and annual ring width was taken into account. Tangential cell width was not significantly affected by treatment at Flakaliden. At Asa fertilization caused a small increase on tangential cell width. Ring width was positively affected by treatment and is an important factor explaining the effects on primarily cell wall thickness and radial cell width.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: christina.lundgren@spm.slu.se (email)
article id 438, category Research article
Christina Lundgren. (2004). Cell wall thickness and tangential and radial cell diameter of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 438. https://doi.org/10.14214/sf.438
Two Norway spruce nutrient trials were used to evaluate the effects of fertilization and irrigation on transverse tracheid dimensions. Three different treatments and a control (C) were used; daily irrigation (I), daily liquid fertilization (IL) and an annual solid fertilization (F). The nutrient optimisation was based on foliage analysis and both liquid and solid fertilization essentially comprised the same amount of nutrients but the latter was applied annually in solid form. The cell measurements; cell wall thickness, radial and tangential cell widths, were obtained using image analysis (SilviScan at CSIRO, Melbourne, Australia). Mean annual cell wall thickness was decreased by fertilization (F and IL) on both sites whereas no effect of the irrigation on wall thickness could be detected. Radial cell width was increased by treatment at Flakaliden but at Asa the effect of irrigation and fertilization was reversed when the data structure i.e. development from pith and out and annual ring width was taken into account. Tangential cell width was not significantly affected by treatment at Flakaliden. At Asa fertilization caused a small increase on tangential cell width. Ring width was positively affected by treatment and is an important factor explaining the effects on primarily cell wall thickness and radial cell width.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: christina.lundgren@spm.slu.se (email)
article id 438, category Research article
Christina Lundgren. (2004). Cell wall thickness and tangential and radial cell diameter of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 438. https://doi.org/10.14214/sf.438
Two Norway spruce nutrient trials were used to evaluate the effects of fertilization and irrigation on transverse tracheid dimensions. Three different treatments and a control (C) were used; daily irrigation (I), daily liquid fertilization (IL) and an annual solid fertilization (F). The nutrient optimisation was based on foliage analysis and both liquid and solid fertilization essentially comprised the same amount of nutrients but the latter was applied annually in solid form. The cell measurements; cell wall thickness, radial and tangential cell widths, were obtained using image analysis (SilviScan at CSIRO, Melbourne, Australia). Mean annual cell wall thickness was decreased by fertilization (F and IL) on both sites whereas no effect of the irrigation on wall thickness could be detected. Radial cell width was increased by treatment at Flakaliden but at Asa the effect of irrigation and fertilization was reversed when the data structure i.e. development from pith and out and annual ring width was taken into account. Tangential cell width was not significantly affected by treatment at Flakaliden. At Asa fertilization caused a small increase on tangential cell width. Ring width was positively affected by treatment and is an important factor explaining the effects on primarily cell wall thickness and radial cell width.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: christina.lundgren@spm.slu.se (email)

Category: Article

article id 7291, category Article
Paul Wallden. (1934). Tutkimuksia koivupuun anatoomisen rakenteen ja teknillisten ominaisuuksien keskinäisestä riippuvuudesta solumittauksien perusteella. Acta Forestalia Fennica vol. 40 no. 14 article id 7291. https://doi.org/10.14214/aff.7291
English title: Studies on relationship between anatomical structure and technical properties of birch wood.

Birch wood is used widely in wooden structures where mechanical strength is needed. The aim of the research was to study the influence of the relative share of mechanically weak tracheids, and length of the wood fibers on specific gravity and bending strength of downy birch (Betula pubescens Ehrh.) wood. According to the results, the strength of wood is strongly dependent on the relative share of tracheids, and length of the libriform cells. The strength of the wood increases when the share of tracheids decreases and the length of libriform cells increases. The specific gravity can be used as an indication of the strength of wood, especially if it is possible to analyze the structure of the wood.

The PDF includes a summary in German.

  • Wallden, ORCID ID:E-mail:
article id 5483, category Article
Erkki Verkasalo. (1992). Relationships of the modulus of elasticity and the structure of Finnish Scots pine wood. Silva Fennica vol. 26 no. 3 article id 5483. https://doi.org/10.14214/sf.a15644

The paper presents preliminary results on the relationships of the longitudinal modulus of elasticity (E) in bending, based on ISO Standard 3349 tests on small, clear specimens, and some basic characteristics of Finnish Scots pine (Pinus sylvestris L.) wood. A manual image analysis method – quantitative stereological counting – was introduced and applied for the investigations of wood structure.

The main results were consistent with those from the prior research. The range of E was 9.7 to 19.1 GPa. Increase in especially fibre density index (R2 = 0.95), weight density and specific gravity (R2 = 0.90), Runkel’s ratio, coefficient of cell rigidity and number of growth rings per cross-sectional unit area, but also in latewood percentage (R2 = 0.58) resulted in an increase in E. Increase in growth ring width, particularly in the width of the late wood section within a ring (R2 = 0.63 to 0.90) had a reverse effect. Cell wall thickness did not show any clear effect. Except for tracheid diameter, the relationships were stronger for the variables determined in the tangential than in the radial wood direction.

Quantitative stereological counting has been used to some degree in the Finnish wood research. The procedure is technically feasible and easy to use. A large sample of counting areas is frequently needed to obtain accurate mean results for the size and distribution of the features. Because the actual analysis points are located at a fixed distance from each other, the method is not in principle well suited for wood with a regular and simple structure, as Scots pine. However, the good correlations between E and some characteristics obtained with stereological counting did not support this misgiving.

The PDF includes an abstract in Finnish.

  • Verkasalo, ORCID ID:E-mail:
article id 5371, category Article
Riitta Laurila. (1989). Pieniläpimittaisen männyn kuituominaisuudet. Silva Fennica vol. 23 no. 1 article id 5371. https://doi.org/10.14214/sf.a15530
English title: Fibre properties in Pinus sylvestris pulpwood.

Properties of fibres in pulpwood, especially length, width and the thickness of walls in tracheids, are essential for strength properties of pulp and paper. Length and width of tracheids increase from pith to surface in radial direction. Young and small-sized stems have also smaller fibres. Small-sized Pinus sylvestris L. test trees had tracheids that were shorter both in stems and knot wood than those in normal sized trees. However, cell walls in test trees were as thick as in normal sized trees. It seems that especially the L/T -ratio (length/thickness) in small stems is worse than in normal sized pulp wood.

The PDF includes an abstract in English.

  • Laurila, ORCID ID:E-mail:
article id 5228, category Article
Pekka Saranpää. (1985). Kontortamännyn runkopuun trakeidien pituuden, halkaisijan ja soluseinän paksuuden vaihtelu. Silva Fennica vol. 19 no. 1 article id 5228. https://doi.org/10.14214/sf.a15407
English title: Length, diameter and cell wall thickness of tracheids in mature lodgepole pine bole wood.

Variation in tracheid morphology were examined for the bole wood of lodgepole pine (Pinus contorta Douglas ex Loudon) grown in Southern and Central Finland. Tracheid lengths were examined in a fast-grown and in slow-grown trees from three stands. Tracheid length increased with increasing height to 4–8 metres and decreased after that, and increased also with increasing age from the pith. The variation between stems was high. The shortest tracheids were about 1.11 mm near the piths and the longest about 4.10 mm near the bark.

Tracheid diameter and cell wall thickness were measured for the total number of 16 stems from Southern and Central Finland. Tracheid diameter increased with increasing distance from pith and the largest tracheids were at a height of 4–8 metres. Cell wall thickness varied independently of height in the stem. Summerwood cell wall thickness was twice that of springwood. There was a difference of 0.6 μm in springwood and 1.0 μm in summerwood double cell wall thickness between the two stands. Cell wall percentage was 29±4.7 in springwood and 69±7.3 in summerwood.

The PDF includes a summary in English.

  • Saranpää, ORCID ID:E-mail:

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