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Articles containing the keyword 'trees'.

Category: Research article

article id 10373, category Research article
Chintan Sheth, Aparajita Datta, Devathi Parashuram. (2020). Persistent loss of biologically-rich tropical forests in the Indian Eastern Himalaya. Silva Fennica vol. 54 no. 3 article id 10373. https://doi.org/10.14214/sf.10373
Highlights: We found a high rate of deforestation occurring in a state managed reserve forest in Indian Eastern Himalaya; Fine-scale analysis showed considerable forest loss around nesting trees for hornbills; Forest monitoring, protection and honest governance are required to effectively protect forests in the Eastern Himalaya biodiversity hotspot.

Using fine-resolution satellite imagery from multiple satellite data products, we assessed the change in forest cover of a state-managed Reserve Forest (RF) located in India’s Eastern Himalaya biodiversity hot-spot. 4.6% of forest cover was lost from Papum RF between 2013 and 2017 at the rate of 8.2 km2 year–1. Three species of hornbills: Great Hornbill Buceros bicornis Linnaeus, 1758, Wreathed Hornbill Rhyticeros undulatus (Shaw, 1811) and Oriental Pied Hornbill Anthracoceros albirostris (Shaw, 1808), that are functionally important are found here with nesting habitat in the areas affected by illegal logging. Therefore, we assessed the habitat loss within a 1 km radius around 29 nest trees. From 2011 to 2019, forest cover declined from 38.55 km2 to 21.94 km2 around these hornbill nest trees. Illegal logging is the main driver that is depleting forest cover within this important bird area. Our results highlight the ongoing threats to biologically-rich forests and the need for urgent measures to halt this loss. We suggest that this study has practical implications for the monitoring and governance of state-managed forests in Arunachal Pradesh.

  • Sheth, #4 Ananda Nilaya, 4th Main Road, Siddhivinayaka Layout, Bengaluru 560097, Karnataka, India ORCID ID:E-mail: chintz604@gmail.com
  • Datta, Nature Conservation Foundation, 1311,“Amritha”, 12th Main, Vijayanagar 1st Stage, Mysore 570017, Karnataka, India ORCID ID:E-mail: aparajita@ncf-india.org (email)
  • Parashuram, Nature Conservation Foundation, 1311,“Amritha”, 12th Main, Vijayanagar 1st Stage, Mysore 570017, Karnataka, India; Department of Geography, University of Cambridge, CB2 3EN, United Kingdom ORCID ID:E-mail: dp608@cam.ac.uk
article id 9968, category Research article
Hubert Lachowicz, Anna Bieniasz, Rafał Wojtan. (2019). Variability in the basic density of silver birch wood in Poland. Silva Fennica vol. 53 no. 1 article id 9968. https://doi.org/10.14214/sf.9968
Highlights: Location, tree age and forest habitat type, and the interactions between those factors, have a statistically significant impact on the basic density of silver birch wood; The average basic density of silver birch wood increases with the age of the tree.

This work presents the findings of a study concerning variability in the basic density of silver birch (Betula pendula Roth) wood, depending on the geographical location of tree stands, the age and thickness of the trees, the forest habitat type, and interactions between some of these factors. The study was carried out on wood from trees aged approximately 30, 50 and 70 years in 12 forest districts located throughout Poland. In total 4777 wood samples, taken from 306 trees from 51 test plots, were measured. The location, the age of the trees, the thickness of the trees and the forest habitat type, as well as interactions between these factors, proved to have a significant influence on the basic density of silver birch wood. The highest mean values of the basic density of the birch wood were found in Sokołów forest district on the FBF habitat type (549 kg m–3) and in Giżycko forest district on the FMBF habitat type (548 kg m–3). For the entire set of examined material, the average values of the basic density of wood increase with tree age. For the examined material originating in FBF and FMBF habitats the average values of basic density showed no significant differences; however, in the cases of the forest districts of Giżycko, Łobez and Rudziniec, significant differences in the analysed property were observed.

  • Lachowicz, Department of Forest Utilization, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland ORCID ID:E-mail: hubert.lachowicz@wl.sggw.pl
  • Bieniasz, Department of Forest Utilization, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland ORCID ID:E-mail: anna.bieniasz@wl.sggw.pl (email)
  • Wojtan, Laboratory of Dendrometry and Forest Productivity, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland ORCID ID:E-mail: rwojtan@wl.sggw.pl
article id 1563, category Research article
Kristina Ahnlund Ulvcrona, Dan Bergström, Urban Bergsten. (2017). Stand structure after thinning in 1–2 m wide corridors in young dense stands. Silva Fennica vol. 51 no. 3 article id 1563. https://doi.org/10.14214/sf.1563
Highlights: Boom corridor thinning (BCT) results in more stand structure heterogeneity than conventional thinning or pre-commercial thinning (PCT), maintaining both smaller-diameter trees and deciduous species; Neither dominant height nor number of possible future crop trees is jeopardized, and boom corridor thinning results in higher values of stem volume and biomass; The technique is flexible as various corridor types give similar stand structure results.

Boom corridor thinning (BCT) has been proposed as a cost-effective technique for biomass thinning (BT) in young dense stands. The objective of this study was to determine how various BCT operations affect stand structure following biomass thinning and to compare the results with conventional selective thinning methods. Two series of field experiments were established; BCT 1-series: Three sites in south of Sweden (9 and 11 m in mean and dominating tree height) with five treatments, including a control, conventional selective thinning and three BCT treatments (1 m and 2 m wide corridors and selective BCT). The second BCT series: Three regions in Sweden (in the north, centre and in the south), with two stand sites in each region with different tree heights (4/9 m and 5/10 m in mean/dominating tree height). Treatments were control, pre-commercial thinning (PCT), conventional selective thinning and BCT (high and low thinning). Following the first biomass thinning, BCT regimes and selective thinning methods resulted in similar stand structures based on the number of possible future crop trees (>80 mm in diameter at breast height). However, BCT maintained a higher diversity of tree sizes as well as more stems per hectare, including deciduous species, than the selective thinning approaches. The stands after BCT should have more vertical complexity, especially when compared to pre-commercial thinning. The structural heterogeneity resulting from BCT may also increase stand biodiversity and ecosystem service values.

  • Ahnlund Ulvcrona, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: kristina.ulvcrona@slu.se (email)
  • Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: dan.bergstrom@slu.se
  • Bergsten, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: urban.bergsten@slu.se
article id 892, category Research article
M. Carmen San José, Laura V. Janeiro, Elena Corredoira. (2013). Micropropagation of threatened black alder. Silva Fennica vol. 47 no. 1 article id 892. https://doi.org/10.14214/sf.892
Micropropagation techniques are valuable tools for propagating, conserving and restoring trees. An efficient micropropagation method involving axillary shoot proliferation of material obtained from mature European alder (Alnus glutinosa (L.) Gaertn.) trees was developed. Branch segments from trees aged 20–30 years were forced to flush, and explants derived from new shoots were cultured on Woody Plant Medium supplemented with 8.88 µM benzyladenine and 2.85µM indole-3-acetic-acid. In vitro establishment was achieved in all five genotypes evaluated. Shoot cultures were maintained by sequential subculture of explants on the same medium supplemented with 0.88–0.44 µM benzyladenine and 2.85 µM indole-3-acetic acid. Transfer to fresh medium every 3 weeks during a 9-week multiplication period and the inclusion of 2.28 µM zeatin during the last 3 weeks of culture improved the multiplication rate and shoot quality. Use of 2% glucose as the carbohydrate source produced better results than 3% sucrose for shoot proliferation. In vitro rooting of shoots was achieved with 2% glucose and 0.49 µM indole-3-butyric acid for 7 days, followed by in vitro culture on auxin-free medium for 21 days. Rooted plantlets were acclimatized to the greenhouse and were viable for reintroduction into the natural habitat.
  • San José, Instituto de Investigaciones Agrobiológicas de Galicia (IIAG), CSIC, Apartado 122, 15080 Santiago de Compostela, Spain ORCID ID:E-mail: sanjose@iiag.csic.es (email)
  • Janeiro, INLUDES, Diputación Provincial de Lugo, Ronda de la Muralla 140, 27004 Lugo, Spain ORCID ID:E-mail: lauravj68@hotmail.com
  • Corredoira, Instituto de Investigaciones Agrobiológicas de Galicia (IIAG), CSIC, Apartado 122, 15080 Santiago de Compostela, Spain ORCID ID:E-mail: elenac@iiag.csic.es
article id 916, category Research article
M. Carmen San José, Lourdes Romero, Laura V. Janeiro. (2012). Effect of indole-3-butyric acid on root formation in Alnus glutinosa microcuttings. Silva Fennica vol. 46 no. 5 article id 916. https://doi.org/10.14214/sf.916
A study of the in vitro rooting process in mature alder (Alnus glutinosa (L.) Gaertn.) shoots is described. Microcuttings from shoots cultured in vitro were transferred to a half-strength Woody Plant Medium containing 0 or 0.1 mg l–1 indole-3-butyric acid (IBA) for 0 to 7 days. The presence of IBA in the medium increased the rooting percentage, number of roots, percentage of lateral roots, and length of the shoots. Histological studies were carried out with shoots treated with 0 or 0.1 mg l–1 IBA for 7 days. According to these criteria, treatment with IBA for 2–3 days proved to be the most successful. In both treatments, substancial reactivation of cell division was observed at the base of the shoots after 1 day. Some cambial zone and adjacent phloem cells became dense cytoplasm, having nuclei with prominent nucleoli. The first cell divisions were also observed at this time. In the treatment with IBA (0.1 mg l–1 for 7 days), meristemoids became individualized, consisting of densely staining cells, by day 3. Identifiable conical shaped root primordia with several cell layers were visible after 4–5 days. Roots with an organized tissue system emerged from the stem after 6 days in the IBA-treated shoots. Meristemoid formation was delayed until the fourth day and root emergence until the eight day in the control treatment (no IBA).
  • San José, Instituto de Investigaciones Agrobiológicas de Galicia, CSIC, Avda de Vigo s/n, Apartado 122, 15780 Santiago de Compostela, Spain ORCID ID:E-mail: sanjose@iiag.csic.es (email)
  • Romero, CIFP Politécnico de Santiago, Santiago de Compostela, Spain ORCID ID:E-mail:
  • Janeiro, INLUDES, Diputación Provincial de Lugo, Lugo, Spain ORCID ID:E-mail: lauravj68@hotmail.com
article id 45, category Research article
Guolei Li, Yong Liu, Yan Zhu, Qing Mei Li, R. Karsten Dumroese. (2012). Effect of fall-applied nitrogen on growth, nitrogen storage and frost hardiness of bareroot Larix olgensis seedlings. Silva Fennica vol. 46 no. 3 article id 45. https://doi.org/10.14214/sf.45
Nursery response of evergreen trees to fall fertilization has been studied widely, but little attention has been given to deciduous trees. Bareroot Olga Bay larch (Larix olgensis Henry) seedlings were fertilized in the nursery with urea at four rates (0, 30, 60, 90 kg N ha–1), with half of each rate applied on two dates (September 16 and October 1, 2009). The seedlings were excavated for evaluation on October 15. In the unfertilized (control) treatment, root and shoot dry mass increased by 100% and 57% respectively, while N concentration in the roots and shoots increased by 43% and 40% during the 30 day period. This indicated that substantial biomass growth during this period did not lead to internal nutrient dilution. Root dry mass increased when fall fertilization rates were ≥ 60 kg N ha–1. Fall fertilization increased N concentrations in root tissue by 48–73%. Compared with the control, shoot tissues of fall fertilized seedlings had slightly higher N concentration and content and significantly higher frost hardiness.
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail: lyong@bjfu.edu.cn (email)
  • Zhu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Li, Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China ORCID ID:E-mail:
  • Dumroese, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA ORCID ID:E-mail:
article id 78, category Research article
Kris Vandekerkhove, Luc De Keersmaeker, Ruben Walleyn, Frank Köhler, Luc Crevecoeur, Leen Govaere, Arno Thomaes, Kris Verheyen. (2011). Reappearance of old-growth elements in lowland woodlands in northern Belgium: Do the associated species follow? Silva Fennica vol. 45 no. 5 article id 78. https://doi.org/10.14214/sf.78
The forest cover of the western European lowland plain has been very low for centuries. Remaining forests were intensively managed, and old-growth elements like veteran trees and coarse woody debris became virtually absent. Only over the last decades have these old-growth elements progressively redeveloped in parks, lanes and forests, and have now reached their highest level over the last 500–1000 years. Biodiversity associated with these old-growth elements makes up an important part of overall forest biodiversity. The ability of species to recolonise the newly available habitat is strongly determined by limitations in their dispersal and establishment. We analyse the current status and development of old-growth elements in Flanders (northern Belgium) and the process of recolonisation by means of specific cases, focussing on saproxylic fungi and saproxylic beetles. Our results show that ‘hotspots’ of secondary old growth, even isolated small patches, may have more potential for specialised biodiversity than expected, and may provide important new strongholds for recovery and recolonisation of an important share of old-growth related species.
  • Vandekerkhove, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail: kris.vandekerkhove@inbo.be (email)
  • De Keersmaeker, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Walleyn, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Köhler, Koleopterologisches Forschungsbüro, Bornheim, Germany ORCID ID:E-mail:
  • Crevecoeur, Genk, Belgium ORCID ID:E-mail:
  • Govaere, Agency of Nature and Forests, Brussels, Belgium ORCID ID:E-mail:
  • Thomaes, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Verheyen, Ghent University, Laboratory of Forestry, Gontrode, Belgium ORCID ID:E-mail:
article id 143, category Research article
Juha Laitila, Jani Heikkilä, Perttu Anttila. (2010). Harvesting alternatives, accumulation and procurement cost of small-diameter thinning wood for fuel in Central Finland. Silva Fennica vol. 44 no. 3 article id 143. https://doi.org/10.14214/sf.143
This study compared harvesting alternatives, accumulation and procurement costs of small-diameter thinning wood chips for fuel, when trees were harvested either as delimbed stemwood or whole trees. The calculation was made for a hypothetical plant located in Central Finland and the radius of the procurement area was 100 km via the existing road network. Cutting was done with conventional harvester head equipped with multi-tree-handling (MTH) accessories, with the logged trees being chipped at the roadside storage. The cost of delimbed stemwood chips at heating plant was 24% higher compared to the cost of whole tree chips. The availability analysis attested that delimbing lowered the regional cutting removal by 42% compared to the whole tree harvesting, when the minimum accumulation for the fuel fraction at the stand was set at 25 m3/ha. Delimbing diminishes the recovery rate at the site, resulting in a diminishing number of potential recovery sites meeting the threshold volume. However, the study showed that the forest energy potential is increased and procurement costs are reduced, if delimbed stemwood is harvested from stands where the whole tree harvesting is not acceptable due to nutrient loss or for other ecological reasons. Intelligent selection of cutting methods for different stands enables minimization of transport distance and control of procurement cost.
  • Laitila, The Finnish Forest Research Institute, Joensuu, Finland ORCID ID:E-mail: juha.laitila@metla.fi (email)
  • Heikkilä, L&T Biowatti Oy, Seinäjoki, Finland ORCID ID:E-mail:
  • Anttila, The Finnish Forest Research Institute, Joensuu, Finland ORCID ID:E-mail:
article id 317, category Research article
Erik Eriksson, Tord Johansson. (2006). Effects of rotation period on biomass production and atmospheric CO2 emissions from broadleaved stands growing on abandoned farmland. Silva Fennica vol. 40 no. 4 article id 317. https://doi.org/10.14214/sf.317
The growth rates and carbon stocks of unthinned young and mature stands of broadleaved trees growing on abandoned farmland were determined to assess whether their management regimes should involve short (15-year) or long (45-year) rotations to maximize biomass production and reductions of CO2 emissions. Dry mass production and mean annual increment (MAI) were calculated for 28 young stands and 65 mature stands of European aspen (Populus tremula L.), common alder (Alnus glutinosa (L.) Gaertn.), grey alder (Alnus incana (L.) Moench.), silver birch (Betula pendula Roth) and downy birch (Betula pubescens Ehrh.) ranging in latitude from 57° to 63° N in Sweden. The potential for using biomass from the stands to replace coal as a fuel and to store carbon was then evaluated both in short and long rotation scenarios. The results indicate that long rotations are beneficial if the objective is to maximize the average carbon stock in biomass. If, on the other hand, the intention is to optimize reductions in atmospheric CO2 emissions, rotations should be short for aspen, silver birch and grey alder stands. For downy birch and common alder, the MAI was higher for the mature stands than the young stands, indicating that in these species the mature stands are superior for both storing carbon and replacing fossil fuel. Stands of broadleaved trees grown to produce biofuel on abandoned farmland should be established on fertile soils to promote high MAI. If the MAI is low, the rotation period should be long to maximize the average carbon stock.
  • Eriksson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden ORCID ID:E-mail:
  • Johansson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden ORCID ID:E-mail:
article id 350, category Research article
Ehsan Sayyad, Seyed Mohsen Hosseini, Jamshid Mokhtari, Reza Mahdavi, Seyed Gholamali Jalali, Moslem Akbarinia, Masoud Tabari. (2006). Comparison of growth, nutrition and soil properties of pure and mixed stands of Populus deltoides and Alnus subcordata. Silva Fennica vol. 40 no. 1 article id 350. https://doi.org/10.14214/sf.350
Concerns about decline in soil fertility and long-term productivity of fast-growing plantations have promoted interest in using nitrogen-fixing trees in mixed species plantations. Populus deltoides and Alnus subcordata were planted in five proportions (100P, 67P:33A, 50P:50A, 33P:67A, 100A) in Noor, Iran. After 7 years, the effects of species interactions on tree growth and nutrient concentration in live and senescent leaves and soil properties were assessed. Diameter at breast height and total height of individual Populus trees were positively affected by the presence of Alnus. Nitrogen concentrations in fully expanded and senescent leaves of Populus were higher in mixed plantations than monoculture plantations. The results of nutrition and nutrient return and growth indicated that mixed plantations of these two species were more productive and sustainable than their monoculture plantations. Within the framework of this experiment, it appeared that production was maximized when these two species were grown together in the relative proportions of 50% Populus and 50% Alnus.
  • Sayyad, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
  • Hosseini, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail: hosseini@europe.com (email)
  • Mokhtari, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
  • Mahdavi, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
  • Jalali, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
  • Akbarinia, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
  • Tabari, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran ORCID ID:E-mail:
article id 479, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Ivar Gjerde, Vegard S. Gundersen. (2005). Historical logging, productivity, and structural characteristics of boreal coniferous forests in Norway. Silva Fennica vol. 39 no. 3 article id 479. https://doi.org/10.14214/sf.479
Conservation of forest biodiversity has brought about an interest in evaluating the naturalness of forests, and to locate and protect semi-natural and old-growth forests in the Fennoscandian countries. However, it is not always clear how natural these forests really are, and how the past management history has affected their present structural composition. We studied the relationships between cut stumps from historical logging activity (50–100 years ago) and forest structural characteristics of today in a total of 385 0.25 ha plots in three boreal coniferous forests which are parts of National Nature Reserves in Norway. We also studied how forest productivity influenced these relationships. In plots with negligible logging impact we found the amount of living trees, dead wood, and size of the oldest trees mainly to increase with increasing productivity, whereas the age of the oldest trees decreased. The amount of deciduous trees was generally low irrespective of productivity. The intensity of logging did not consistently influence most of these forest structural variables, neither at low- nor at high-productive sites. The only consistent relationship in all study areas was a decreasing amount of dead wood with increasing logging intensity at high-productive sites. Also, the decay class distribution of dead wood was more right-skewed (indicating on-going accumulation of dead wood) the more logging had occurred at high-productive sites. Except from the effects on dead wood, previous logging does not show up as a major determinant of other stand structures of today.
  • Storaunet, Norwegian Forest Research Institute, Høgskolevegen 8, NO-1432 Ås, Norway ORCID ID:E-mail: ken.storaunet@skogforsk.no (email)
  • Rolstad, Norwegian Forest Research Institute, Høgskolevegen 8, NO-1432 Ås, Norway ORCID ID:E-mail:
  • Gjerde, Norwegian Forest Research Institute, Fanaflaten 4, NO-5244 Fana, Norway ORCID ID:E-mail:
  • Gundersen, Norwegian Forest Research Institute, Fanaflaten 4, NO-5244 Fana, Norway ORCID ID:E-mail:
article id 507, category Research article
Tysk Staffan Ericsson, Lars Östlund, Rikard Andersson. (2003). Destroying a path to the past – the loss of culturally scarred trees and change in forest structure along Allmunvägen, in mid-west boreal Sweden. Silva Fennica vol. 37 no. 2 article id 507. https://doi.org/10.14214/sf.507
The tradition to blaze trees to mark trails and boundaries is very old in northern Scandinavia. The disappearance of culturally modified trees (i.e. trees with trail blazes) and changes in forest structure along a section of an old bridle trail in boreal Sweden was analyzed using historical maps and forest surveys from the period 1876 to the year 2000. Remaining blazed trees were located during a field study and selected scars were dated. In total 104 scarred living and dead trees were found. The scars originated from the early 1500s to the early 1900s. Analysis of the forest surveys showed that the forest along the trail was dominated by older trees, and that the majority of the scarred trees probably were present, throughout the 19th century. By the mid 20th century logging had begun to affect the tree age along the trail and in 1974 no stands older than 180 years were present. A conservative estimate shows that around 90% of the original blazed trees have vanished. The trail was interpreted as have being lined for centuries with scarred trees which gradually have been destroyed during the 20th century. Culturally modified trees constitute an unique source of information for understanding pattern of old trails as well as of past human land use and movement in the landscape prior to the 20th century. This biological archive have to a large extent been destroyed by forestry activities and it is therefore very important to survey, recount and protect the trees that are still present.
  • Ericsson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail: staffan@delta.se (email)
  • Östlund, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Andersson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail:
article id 507, category Research article
Tysk Staffan Ericsson, Lars Östlund, Rikard Andersson. (2003). Destroying a path to the past – the loss of culturally scarred trees and change in forest structure along Allmunvägen, in mid-west boreal Sweden. Silva Fennica vol. 37 no. 2 article id 507. https://doi.org/10.14214/sf.507
The tradition to blaze trees to mark trails and boundaries is very old in northern Scandinavia. The disappearance of culturally modified trees (i.e. trees with trail blazes) and changes in forest structure along a section of an old bridle trail in boreal Sweden was analyzed using historical maps and forest surveys from the period 1876 to the year 2000. Remaining blazed trees were located during a field study and selected scars were dated. In total 104 scarred living and dead trees were found. The scars originated from the early 1500s to the early 1900s. Analysis of the forest surveys showed that the forest along the trail was dominated by older trees, and that the majority of the scarred trees probably were present, throughout the 19th century. By the mid 20th century logging had begun to affect the tree age along the trail and in 1974 no stands older than 180 years were present. A conservative estimate shows that around 90% of the original blazed trees have vanished. The trail was interpreted as have being lined for centuries with scarred trees which gradually have been destroyed during the 20th century. Culturally modified trees constitute an unique source of information for understanding pattern of old trails as well as of past human land use and movement in the landscape prior to the 20th century. This biological archive have to a large extent been destroyed by forestry activities and it is therefore very important to survey, recount and protect the trees that are still present.
  • Ericsson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail: staffan@delta.se (email)
  • Östlund, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Andersson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden ORCID ID:E-mail:
article id 675, category Research article
Per Linder, Peter Jonsson, Mats Niklasson. (1998). Tree mortality after prescribed burning in an old-growth Scots pine forest in northern Sweden. Silva Fennica vol. 32 no. 4 article id 675. https://doi.org/10.14214/sf.675
Tree mortality and input of dead trees were studied after a prescribed burning in a forest reserve in northern Sweden. The stand was a multi-layered old-growth forest. The overstorey was dominated by Scots pine (Pinus sylvestris L.) and the understorey consisted of mixed Scots pine and Norway spruce (Picea abies L. Karst.). Ground vegetation was dominated by ericaceous dwarf-shrubs and feathermosses. The stand has been affected by six forest fires during the last 500 years. The prescribed burning was a low intensity surface fire that scorched almost 90% of the ground. Tree mortality for smaller pines and spruces (DBH < 10 cm) was over 80% in the burned parts of the reserve. For larger pines, 10–50 cm DBH, mortality showed a decreasing trend with increasing diameter, from 14% in class 10–20 cm DBH to 1.4% in class 40–50 cm DBH. However, pines with DBH ≥ 50 cm had a significantly higher mortality, 20%, since a high proportion of them had open fire scars containing cavities, caused by fungi and insects, which enabled the fire to burn inside the trunks and hollow them out. The fire-induced mortality resulted in a 21 m3 ha–1 input of dead trees, of which 12 m3 ha–1 consisted of trees with DBH ≥ 30 cm. An increased mortality among larger trees after low-intensity fires has not previously been described in Fennoscandian boreal forests, probably owing to a lack of recent fires in old-growth stands. However, since large pines with open fire scars were once a common feature in the natural boreal forest, we suggest that this type of tree mortality should be mimicked in forestry practices aiming to maintain and restore natural forest biodiversity.
  • Linder, Department of Forest Vegetation Ecology, SLU, S-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Jonsson, Department of Forest Vegetation Ecology, SLU, S-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Niklasson, Southern Swedish Forest Research Centre, SLU, Box 49, S-230 53 Alnarp, Sweden ORCID ID:E-mail:

Category: Review article

article id 535, category Review article
Thomas J. Givnish. (2002). Adaptive significance of evergreen vs. deciduous leaves: solving the triple paradox. Silva Fennica vol. 36 no. 3 article id 535. https://doi.org/10.14214/sf.535
Patterns in the dominance of evergreen vs. deciduous plants have long interested ecologists, biogeographers, and global modellers. But previous models to account for these patterns have significant weaknesses. Bottom-up, mechanistic models – based on physiology, competition, and natural selection – have often been non-quantitative or restricted to a small range of habitats, and almost all have ignored belowground costs and whole-plant integration. Top-down, ecosystem-based models have succeeded in quantitatively reproducing several patterns, but rely partly on empirically derived constants and thresholds that lack a mechanistic explanation. It is generally recognized that seasonal drought can favor deciduous leaves, and that infertile soils can favor long-lived evergreen leaves. But no model has yet explained three great paradoxes, involving dominance by 1) evergreens in highly seasonal, boreal forests, 2) deciduous larch in many nutrient-poor peatlands, and 3) evergreen leaf-exchangers in nutrient-poor subtropical forests, even though they shed their leaves just as frequently as deciduous species. This paper outlines a generalized optimality model to account for these and other patterns in leaf longevity and phenology, based on maximizing whole-plant carbon gain or height growth, and building on recent advances in our understanding of the quantitative relationships of leaf photosynthesis, nitrogen content, and mass per unit area to leaf life-span. Only a whole-plant approach can explain evergreen dominance under realistic ecological conditions, or account for the boreal paradox, the larch paradox, the leaf-exchanger paradox, and expected shifts in shade tolerance associated with leaf phenology. Poor soils favor evergreens not merely by increasing the costs of nutrient acquisition, but also by depressing the maximum rate of photosynthesis and thus the seasonal contrast in photosynthetic return between leaves adapted to favorable vs. unfavorable conditions. The dominance of evergreens in western North America beyond the coastal zone of mild winters and winter rainfall appears related to the unusually long photosynthetic season for evergreen vs. deciduous plants there. Future models for optimal leaf phenology must incorporate differences between evergreen and deciduous plants in allocation to photosynthetic vs. non-photosynthetic tissue, rooting depth, stem allometry, xylem anatomy, and exposure to herbivores and leaching, and analyze how these differences interact with the photosynthetic rate, transpiration, and nutrient demands of leaves with different life-spans to affect rates of height growth in specific microsites.
  • Givnish, Dept of Botany, University of Wisconsin, Madison, WI 53706, USA ORCID ID:E-mail: givnish@facstaff.wisc.edu (email)

Category: Research note

article id 897, category Research note
Lars Lundqvist, Susanne Spreer, Christer Karlsson. (2013). Volume production in different silvicultural systems for 85 years in a mixed Picea abies–Pinus sylvestris forest in central Sweden. Silva Fennica vol. 47 no. 1 article id 897. https://doi.org/10.14214/sf.897
A long-term comparison of different silvicultural systems was established in 1923 in central Sweden, in an uneven-aged mixed Norway spruce–Scots pine forest (Picea abies (L.) Karst. – Pinus sylvestris L.) with about 85% spruce and 15% pine. The five treatments consisted of two examples of even-aged management 1) clear-cutting followed by planting, and 2) seed tree regeneration, one uneven-aged management 3) selection system, one exploiting treatment 4) diameter limit cut, and 5) one untreated control plot. Each treatment plot was 1 ha, 100 m × 100 m. The plots were measured and managed at irregular intervals, ranging from 7 to 15 years. In 2007–2008 the even-aged treatments and the diameter limit cut were repeated and a new rotation started. Mean annual volume increment during the whole observation period differed widely between the treatments, partly because of differences in species composition over time, with treatment clear-cutting followed by planting at the top, and the control at the bottom. Treatment selection system gave only about 60% of planting, but this was probably largely an effect of too small growing stock during the first roughly 50 years. When the growing stock was increased, periodic annual volume increment increased to about 80% of the mean annual volume increment in the even-aged, planted plot.
  • Lundqvist, Deparment of Forest Ecology and Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden ORCID ID:E-mail: lars.lundqvist@slu.se (email)
  • Spreer, Sveaskog Förvaltnings AB, Ljusdal, Sweden ORCID ID:E-mail: susanne.spreer@sveaskog.se
  • Karlsson, Field Research Unit, Swedish University of Agricultural Sciences, Siljansfors, Sweden ORCID ID:E-mail: christer.karlsson@slu.se

Category: Article

article id 7158, category Article
Olavi Huuri. (1965). Männyn- ja kuusenkäpyjen varastoinnin vaikutus niistä saatavan siemenen itävyyteen. Acta Forestalia Fennica vol. 78 no. 5 article id 7158. https://doi.org/10.14214/aff.7158
English title: The effects of storage in cones on the viability of pine and spruce seeds.

Seed storing experiments with cones of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were conducted in Oitti seed extracting plant in Southern Finland from February to December 1955. The pine cones were stores for 267 and the spruce coned for 304 days. In four of the storage methods the cones were packed in sacks and another four in wooden boxes. Sample of cones were taken once a month, seeds were extracted and the germinative capacity was tested. The remaining extracted seeds were placed in storage, and in January 1956 moved to cold seed cellar until 1962, when the viability of the seeds was tested.

According to the results, cleaned pine cones can be stores for at least nine months using almost all methods of storage which are commonly used at our seed traction plants, without hazarding the usability of the seeds. The seeds in spruce cones, however, seemed to be more sensitive to conditions during the storage. The germinative capacity of the spruce seeds began to decrease after the beginning of May. Later the seeds were infected with mould, which increased towards the end of the experiment.

Thus, preservation of the germinative capacity of the seeds of pine and spruce requires storage in different conditions. The results suggest that extraction of spruce seeds should be finished during the cold winter months. It seems that seed in the cones of pine and spruce endure storage in piles of paper or cloth sacks at least as well as in wooden boxes. Occasional warming of the storage, snow and foreign material among the cones and an over meter thick cone layer decreased the germinative capacity of spruce seeds during spring and summer. Spruce seeds that had been extracted immediately after collecting of the cones preserved their germinative capacity well during an eight years storage period.

The PDF includes a summary in English.

  • Huuri, ORCID ID:E-mail:
article id 7114, category Article
Paavo Yli-Vakkuri. (1960). Metsiköiden routa- ja lumisuhteista. Acta Forestalia Fennica vol. 71 no. 5 article id 7114. https://doi.org/10.14214/aff.7114
English title: Snow cover and ground frost in Finnish forests.

Snow cover and ground frost was studied in 29 forest stands in Southern and Central Finland in 1957–1959. The tree species influenced greatly accumulation of snow on the forest floor. Norway spruce (Picea abies (L.) Karst.) retains snow in its crown. In addition, snow and water falling from the branches compress the snow cover under the trees, and the ground freezes deeper because of the shallow snow cover. In the spring, the dense crown prevents rain and radiation reaching the ground, which remains cold longer. However, ground frost may protect spruce, which has a weak root system, from wind damages.

Scots pine (Pinus sylvestris L.) has similar, but milder, effects on snow cover within the forest. The crowns of pine seedlings and young trees pass snow easily, but later the crowns intercept it considerably. The lower branches are, however, high up and the snow is evenly spread on the ground. The deciduous trees intercept little snow and in the spring the snow smelts and the frozen soil thaws early. The snow conditions of deciduous forests are, however, changed by a spruce undergrowth.

It can be assumed that the unfavourable conditions in spruce forests can be alleviated by thinning. Also, mixture of pine and deciduous trees can transform the conditions more favourable in the spruce stands.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7486, category Article
Paavo Yli-Vakkuri. (1959). Siemensiipien hankaajista ja niiden vaikutuksesta siemenen itävyyteen. Acta Forestalia Fennica vol. 68 no. 4 article id 7486. https://doi.org/10.14214/aff.7486
English title: On machines for abrading seed wings and their influence on the germinative capacity of the seed.

This paper deals with two machines designed for abrading seed wings, and their influence on the germinative capacity of seed of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.). Both machines are commonly used in Finland.

The results of the study indicate that the act of abrading may cause slight or even serious injuries to the seed. Slight injuries of about 3% are probably not easily avoided if mechanical abrading is resorted to. It must be noted, however, that even this reduction in germinative capacity causes significant yearly loss. If the reduction in germinative capacity is greater, which seems to be possible, it is advisable to test the mechanism of the machine and its method of abrading. As the clearance of the machines can affect the extent of injuries, all machines should be tested. If possible, a continual operation control should be arranged. It could, at the same time, to supply material for improving the abrading method and equipment.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7440, category Article
Erkki K. Kalela. (1954). Mäntysiemenpuiden ja -puustojen juurisuhteista. Acta Forestalia Fennica vol. 61 no. 28 article id 7440. https://doi.org/10.14214/aff.7440
English title: Root systems of Scots pine seed trees and stands.

Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.

Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Kalela, ORCID ID:E-mail:
article id 7436, category Article
E. Salo. (1954). Puiden teknillinen vikaisuus ja sen vaikutus puuston arvoon. Acta Forestalia Fennica vol. 61 no. 24 article id 7436. https://doi.org/10.14214/aff.7436
English title: Technical defects of trees and their effect upon timber value.

In acquiring land for the population displaced by the Second World War, the forest had to be priced, according to the Land Reclamation Act of 1945, separately for land and timber. Technical defects in the growing stock were to be taken account in the form of a total reduction in the value of the stock. Generally, it was to be 5-15% of the total value. The present investigation aims at checking the reduction percentages.

When the reduction in the felling value of the growing stock caused by the defects is estimated, the reduction is defined for each timber assortment, and the total reduction is calculated from these values. The timber assortments have big variation in prices, therefore defects in the most valuable assortments can have big effect on the total value of the growing stock. According to the study, the decree implementing the Land Reclamation Act did not in some cases allow for price reductions for defects on a sufficiently small scale to correspond to real conditions.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.
The PDF includes a summary in English.

  • Salo, ORCID ID:E-mail:
article id 7370, category Article
Erkki K. Kalela. (1942). Männyn taimien juurien suhtautumisesta emäpuun juuriin. Acta Forestalia Fennica vol. 50 no. 17 article id 7370. https://doi.org/10.14214/aff.7370
English title: Roots of a seedling in relation to roots of the mother tree.

The study is based on observations in a Scots pine (Pinus sylvestris L.) stand on a dry upland forest site in Karhumäki, where a 10-15-year old seedling stand grew under a hold-overs of larger trees that had been left in the site in a previous felling. The root systems of 80-120 cm tall seedlings growing around single mother trees were unearthed. Root maps were drawn of the root systems of 120 seedlings.

No seedlings grew around old, large hold-overs. It seems that seedlings could not compete with their root system. If the hold-overs were stunted in their growth, seedlings grew also under the canopy of the mother tree. 90% of the seedlings had a tap root. Rest of the roots grew horisontally in the topsoil. Around a vigorous mother tree, the seedlings grew their roots away from the mother tree. Hold-overs that had belonged originally to the lower canopy layer of the old forest did not have similar effect on the root orientation of the seedlings. Their roots had been previously affected by trees of higher canopy layer, later removed in the felling.

The PDF includes a summary in German.

  • Kalela, ORCID ID:E-mail:
article id 7358, category Article
Arvo Ylinen. (1942). Über den Einfluss des Spätholzanteils und der Rohwichte auf die Festigkeits- und elastischen Eigenschaften des Nadelholzes. Acta Forestalia Fennica vol. 50 no. 5 article id 7358. https://doi.org/10.14214/aff.7358
English title: The effect of the amount of the summer wood on the elastic and tensile properties of coniferous trees.

The dependence of elastic and tensile properties of coniferous trees on the share of summer wood can be presented as linear functions, if the soft wood is considered as a statically indefinite structure. By eliminating the share of summer wood by a certain function presented in the article, the elastic and tensile properties are the linear functions of density.

The functions are proved right by conducting strength tests on pine. Practical implications of the derived functions in rating the quality of wood are also presented.

The PDF contains a summary in Finnish. 

  • Ylinen, ORCID ID:E-mail:
article id 5629, category Article
Risto Sievänen, Eero Nikinmaa, Jari Perttunen. (1997). Evaluation of importance of sapwood senescence on tree growth using the model Lignum. Silva Fennica vol. 31 no. 3 article id 5629. https://doi.org/10.14214/sf.a8531

The effects of two alternative formulations of sapwood senescence on the behaviour of model LIGNUM (with parameter values adjusted for Scots pine (Pinus sylvestris L.) growing southern Finland) were studied. The two alternatives were autonomous sapwood senescence assuming a maximum age for the tree ring, and sapwood senescence that is controlled by the mortality of foliage. For the latter alternative two hypothetical further mechanisms were stipulated. All the formulations were implemented in LIGNUM. Simulations were made with all model variants for fertile and poor soil conditions using high, normal and low rates of foliage mortality. The simulation results were compared against of a data set consisting of 11 open grown Scots pine trees from southern Finland. Observations of heartwood proportion were used in this study. They show that heartwood starts to increase in trees from age of approximately 20 years onwards. The simulation results showed no differences between fertile and poor soil conditions as regards heartwood formation. Of the variants of foliage-controlled sapwood senescence the one where death of sapwood in a tree segment induces sapwood senescence in the tree parts below only slightly was the best. This and the autonomous sapwood senescence corresponded equally well to the observations. In order to make more refined conclusions additional data and simulations are necessary.

  • Sievänen, ORCID ID:E-mail:
  • Nikinmaa, ORCID ID:E-mail:
  • Perttunen, ORCID ID:E-mail:
article id 5539, category Article
Tore Skrøppa. (1994). Impacts of tree improvement on genetic structure and diversity of planted forests. Silva Fennica vol. 28 no. 4 article id 5539. https://doi.org/10.14214/sf.a9179

After a presentation of basic biodiversity concepts, reviews are made of studies reporting genetic implications of tree improvement activities: seed treatments, seedling production, provenance transfers, plus tree selection, seed production in seed orchards and progeny testing.

Several of the activities may influence the genetic structure and diversity of the planted forests. The general conclusion is, however, that planted forests are at least as genetically diverse as the natural stands that they replace. The diversity in forest management and use is best assurance for the future adaptability of the forests.

  • Skrøppa, ORCID ID:E-mail:
article id 5506, category Article
Risto Heikkilä, Sauli Härkönen. (1993). Moose (Alces alces L.) browsing in young Scots pine stands in relation to the characteristics of their winter habitats. Silva Fennica vol. 27 no. 2 article id 5506. https://doi.org/10.14214/sf.a15667

Moose (Alces alces L.) browsing was studied in young Scots pine (Pinus sylvestris L.) stands mixed with deciduous trees in high-density winter ranges. The proportional use of twig biomass decreased as the availability increased. The total as well as proportional biomass consumption were higher on the moist than on the dry type of forest. The per tree consumption of pine was higher on the moist type, where the availability of pine was lower. Deciduous trees were more consumed on the moist type, where their availability was relatively high. The consumption of pine saplings increased as the availability of birch increased. Pine stem breakages were most numerous when birch occurred as overgrowth above pine and at high birch densities. The availability of other deciduous tree species did not correlate with browsing intensity of Scots pine. Moose browsing had seriously inhibited the development of Scots pines in 6% of the stands, over 60% of available biomass having been removed. Rowan and aspen were commonly over-browsed and their height growth was inhibited, which occurred rarely by birch. There was no difference in the proportion of young stands in forest areas with high and low moose density. A high proportion of peatland forests was found to indicate relatively good feeding habitats in the high-density areas.

The PDF includes an abstract in Finnish.

  • Heikkilä, ORCID ID:E-mail:
  • Härkönen, ORCID ID:E-mail:
article id 5379, category Article
Timo Kuuluvainen, Timo Pukkala. (1989). Effect of Scots pine seed trees on the density of ground vegetation and tree seedlings. Silva Fennica vol. 23 no. 2 article id 5379. https://doi.org/10.14214/sf.a15536

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, ORCID ID:E-mail:
  • Pukkala, ORCID ID:E-mail:
article id 5328, category Article
Max Guignard. (1987). The tree in the Paris and Helsinki areas. Silva Fennica vol. 21 no. 4 article id 5328. https://doi.org/10.14214/sf.a15485

The paper describes some examples from Paris and Helsinki areas, where trees are essential landscape elements. It is typical in France to plant trees around town squares, market places and along streets and roads. In Finland trees are almost always kept close to the house, together with other vegetation protecting the entrance and windows. These traditional uses of trees should be studied to serve landscape management.

The paper is based on a lecture given in the seminar ‘The forest as a Finnish cultural entity’, held in Helsinki in 1986. The PDF includes a summary in Finnish.

  • Guignard, ORCID ID:E-mail:
article id 5256, category Article
Outi Muona, Raimo Hiltunen, Erkki Morén, D. V. Shaw. (1986). Analysis of monoterpene variation in natural stands and plustrees of Pinus sylvestris in Finland. Silva Fennica vol. 20 no. 1 article id 5256. https://doi.org/10.14214/sf.a15435

Variation of monoterpene composition of Pinus sylvestris L. was studied in Southern, Central and Northern Finland using data from both natural stands and plus trees. The natural stands were analysed using different techniques and for fewer terpenes than the plus trees.

There were large differences between areas in the proportion of 3-carene in trees from natural stands, as has been discussed by previous authors. The proportion of 3-carene is bimodally distributed and believed to be controlled by a single gene with large effect. For this reason, we stratified our samples into high carene (>10%) and low carene (<10%) groups. Univariate analysis did not reveal any additional differences between natural populations in different zones for components other than 3-carene. In plus trees, several components showed significant differences, but the proportion of 3-carene did not differ between areas. Multivariate discrimination analysis did not distinguish between areas for natural stands. However, for the plus trees discriminant analysis allowed us to discriminate between the zones relatively efficiently. The proportion of correct classification was greater than 64% using the best methods. The central zone was most distinct, and 80% of its trees were correctly classified. Broad generalizations are not possible due to the limitations imposed by our data. Our analysis of phenotypic variation does not support the suggestion that plus trees selected from the north represent a southern type.

The PDF includes an abstract in Finnish.

  • Muona, ORCID ID:E-mail:
  • Hiltunen, ORCID ID:E-mail:
  • Morén, ORCID ID:E-mail:
  • Shaw, ORCID ID:E-mail:
article id 5133, category Article
F. Scholz. (1981). Genecological aspects of air pollution effects on northern forests. Silva Fennica vol. 15 no. 4 article id 5133. https://doi.org/10.14214/sf.a15360

Natural forest tree populations are adapted to their natural environment. Forest tree species under northern conditions are at the edge of their range where the short growing season and the low winter temperatures are the two main factors limiting their ecological niche. Effects of air pollution on the ecological niche, designated as the environmental conditions that permit a population to survive permanently, are discussed according to G.E. Hutchinson’s concept of the ecological niche. Air pollution as an additional stress factor influences the ecological niche either by the direct influence as an additional dimension of the ecological niche or by interaction with the other dimensions. These interactions are especially important for low level long-term effects of air pollution which can result in reduced resistance to low winter temperature or, due to reduced net assimilation, reduced capability to survive the long period of winter dormancy. These effects influence the boundary of the ecological niche and reduce the area of the biotope of the respective species.

Within the remaining biotope genetic changes in forest tree species take place. Due to individual differences in exposure and susceptibility of trees to air pollution, higher and therefore more exposed trees as well as more susceptible trees will be reduced in reproduction or even be eliminated. This causes genetic changes in the tree population.

  • Scholz, ORCID ID:E-mail:
article id 5059, category Article
Pertti Harstela. (1980). Jäljelle jäävä puusto ja ajouralta toimivat harvennuspuun korjuukoneet. Silva Fennica vol. 14 no. 1 article id 5059. https://doi.org/10.14214/sf.a15000
English title: Remaining trees and machines working from the strip roads in thinning.

In the first part of the study the hindrance of the remaining trees when felling trees by machines working from the strip road in selective thinning was studied on the basis of the literature. In the second part there was geometrically studied the need of schematic thinning in some type stands when bundles are pre-skidded straight-lined to the strip road. In average only 0-1 trees per pre-skidding trail needs to be removed. It was concluded that trees removed from the pre-skidding trail do not significantly increase the need of schematic thinning. Remaining trees do not limit the length of machine booms if the pre-skidding trails are planned during the felling.

The PDF includes a summary in English.

  • Harstela, ORCID ID:E-mail:
article id 5006, category Article
Seppo Kellomäki. (1978). Typpilannoituksen vaikutus havupuiden fotosynteesikapasiteettiin. Silva Fennica vol. 12 no. 3 article id 5006. https://doi.org/10.14214/sf.a14860
English title: Effects of nitrogen fertilization on photosynthetic capacity of coniferous trees.

The effect of nitrogen fertilizers on the photosynthetic capacity of conifers is assessed on the basis of literature. The review emphasizes the role of changes of needle mass as a factor affecting the result of nutrient application. In particular, the increase in needle mass results in a considerable increase in photosynthetic capacity. The effect of fertilization on the photosynthetic rate seems to be of minor importance. The effect on the photosynthetic rate is, however, poorly documented as is the case for the effect of fertilization on the respiration rate. There is evidence that proper application of nitrogen fertilizers may double the photosynthetic capacity of conifers, mainly as a result of increase in needle mass.

The PDF includes a summary in English.

  • Kellomäki, ORCID ID:E-mail:
article id 4931, category Article
Olli Makkonen. (1976). Mitä vanhalla ajalla tiedettiin puiden kasvusta. Silva Fennica vol. 10 no. 1 article id 4931. https://doi.org/10.14214/sf.a14773
English title: What was known in ancient times about growth of trees?
Original keywords: historia; puiden kasvu
English keywords: history; growth of trees

In this article, information about tree growth which was familiar to the learned men in the old days is presented. The time when different tree species start growing, the different growth rate of various tree species, the age of trees, their resistance to injury etc. are discussed.

The PDF includes a summary in English.

  • Makkonen, ORCID ID:E-mail:
article id 4865, category Article
Tauno Kallio. (1972). Esimerkki kuusikon lahovikaisuuden Etelä-Suomessa aiheuttamasta taloudellisesta menetyksestä. Silva Fennica vol. 6 no. 2 article id 4865. https://doi.org/10.14214/sf.a14668
English title: An example on the economic loss caused by decay in growing Norway spruce timber in Southern Finland.

A growing stand of Norway spruce (Picea abies (L.) H. Karst.) marked for cutting was investigated in the winter of 1971–72 in Helsinki in Southern Finland in order to determine the economic loss caused by decay. Taking a sample from growing spruce trees with increment borer is not a reliable method of determining the frequency of decay. The decayed stems were twice measured for assortment cutting into lengths; the first time disregarding the decay and the second time doing the actual assortment cutting according to the grade of timber. The direct economic loss caused by decay was 13% of the price for standing timber. The indirect loss may be as great as the direct loss.

The PDF includes a summary in English.

  • Kallio, ORCID ID:E-mail:
article id 4823, category Article
Christel Palmberg. (1970). Heritabiliteetin arvioiminen eräässä männyn (Pinus silvestris L.) jälkeläiskokeessa. Silva Fennica vol. 4 no. 3 article id 4823. https://doi.org/10.14214/sf.a14612
English title: Estimation of heritability in open-pollinated plus tree progenies of Pinus sylvestris L.

Field experiments of Scots pine (Pinus sylvestris L.) was established by planting seedlings grown from seeds collected from open-pollinated plus trees throughout the country. The 36 progenies represented were planted in 4 blocks as 2+2 transplants in 1960. The main characteristics of the seedlings were measured in 1966 and 1968. Considerable damage had been caused to the stands by moose (Alces alces) and Melampsora pinitorqua Rostr., consequently, therefore, only normally developed seedlings were measured.

Highly significant differences between progenies were found in the number of branches in 1968 and in the ratio of height of tree to the length of the longest branch. In 1968, the differences in height between progenies were not significant, but there were significant differences between blocks both in tree height and length of terminal shoot. Obviously, the edaphic heterogeneity of the site has influenced mainly the juvenile growth of the plants, because in the length of the terminal shoot there could be seen also significant differences between the progenies. There were no significant differences between the progenies in the length of the longest branch, in the angles of the thickest branches, in stem taper and in the diameter of the thickest branch.

The PDF includes a summary in English.

  • Palmberg, ORCID ID:E-mail:
article id 7528, category Article
Einari Vuori. (1913). Die durch Brandkultur entstandenen Nadelholzbestände des Staatsforstes Vesijako. Acta Forestalia Fennica vol. 2 no. 1 article id 7528. https://doi.org/10.14214/aff.7528
English title: Coniferous tree stands of the state forest “Vesijako” reforested through controlled burning.

The study area is state owned forest of Vesijako in southern middle Finland that has earlier been intensively managed with slash-and-burn agriculture. Reforestation of broad-leaved forests into coniferous forest with controlled burning has been studied on 76 sample plots.

The article describes the practice of leasing forest stands to leaseholders who executed the controlled burning and forest regeneration and management according a leasing contract. The results of the reforestation with coniferous trees shows that sowed pine (Pinus silvestris) stands give good results but spruce (Picea abies) must be planted as a seedling.  For the state this method of forest improvement is cost effective  and should be used more widely. 

  • Vuori, ORCID ID:E-mail:
article id 7517, category Article
Juha Nurmi. (1997). Heating values of mature trees. Acta Forestalia Fennica no. 256 article id 7517. https://doi.org/10.14214/aff.7517

The effective heating values of the above and below ground biomass components of mature Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), downy birch (Betula pubescens Ehrh), silver birch (B. pendula Roth), grey alder (Alnus incana (L.) Moench), black alder (A. glutinosa (L.) Gaertn.) and aspen (Populus tremula L.) were studied. Each sample tree was divided into wood, bark and foliage components. Bomb calorimetry was used to determine the calorimetric heating values.

The species is a significant factor in the heating value of individual tree components. The heating value of the wood proper is highest in conifers. Broadleaved species have a higher heating value of bark than conifers. The species factor diminishes when the weighted heating value of crown, whole stems or stump-root-system are considered. The crown material has a higher heating value per unit weight in comparison with fuelwood from small-sized stems or whole trees. The additional advantages of coniferous crown material are that it is non-industrial biomass resource and is readily available. The variability of both the chemical composition and the heating value is small in any given tree component of any species. However, lignin, carbohydrate and extractive content were found to vary from one part of the tree to another and to correlate with the heating value

  • Nurmi, ORCID ID:E-mail:
article id 7682, category Article
Juha Nurmi. (1993). Heating values of the above ground biomass of small-sized trees. Acta Forestalia Fennica no. 236 article id 7682. https://doi.org/10.14214/aff.7682

The heating values of wood, inner and outer bark, and foliage components of seven small-size tree species (Pinus sylvestris L., Picea abies (L.) H. Karst., Betula pubescens Erhr., B. pendula Roth, Alnus incana (L.) Moench, A. glutinosa (L.) Gaertn., Populus tremula L.) were studied. Significant differences were found between species within each component. However, the differences between species for weighted stem, crown and whole-tree biomass are very small. The weighted heating value of the crown mass is slightly higher than that of the stem in all species. The heating value of stem, crown and whole-tree material was found to increase with increasing latitude.

The effective heating value of wood correlated best with the lignin content, inner bark with carbohydrate, and outer bark with carbohydrates and the extractives soluble in alkalic solvents. It is suggested that the determination of the heating value might be used as an indicator of the cellulose content of coniferous wood.

The PDF includes a summary in Finnish.

  • Nurmi, ORCID ID:E-mail:
article id 7670, category Article
Risto Heikkilä. (1991). Moose browsing in a Scots pine plantation mixed with deciduous tree species. Acta Forestalia Fennica no. 224 article id 7670. https://doi.org/10.14214/aff.7670

The utilization of available food resources by the moose (Alces alces L.) was studied in a Scots pine (Pinus sylvestris L.) plantation containing an admixture of deciduous species. Rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) were highly utilized compared to pine and both silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.). However, they were not capable of withstanding continuous browsing by moose owing to their diminished biomass. In total, the browsing intensity (number of browsed twigs/tree) on pine and birch was about double of that on rowan and aspen.

The number of browsed twigs per tree increased as the amount of available main branches increased. The number of bites per available branch, as well as the maximum diameter of the bites, decreased as the density of the plantation increased. Silver birch was more used by moose than pubescent birch as well as planted silver birch compared with naturally regenerated trees.

Main stem breakage was especially common in winter 1988, the average height of the pine and birch trees being over two meters. The tops of broken stems were commonly utilized as food. The increase in moose density and the relatively deep snow cover evidently promoted the incidence of serious damage. The number of undamaged trees/ha was greater in dense than in sparse parts of the stand.

The PDF includes a summary in Finnish.

  • Heikkilä, ORCID ID:E-mail:
article id 7629, category Article
Pekka Kilkki. (1983). Sample trees in timber volume estimation. Acta Forestalia Fennica no. 182 article id 7629. https://doi.org/10.14214/aff.7629

Analysis of sample tree data showed that the major part of the residual variation of the stem volume estimates occurs within the forest stands. The division of the residual variation into the variation within and between populations is the basis for the mean-square error formulae of the volume estimators. Efficiency of different sample tree measurement combinations has been studied by comparing the errors of the volume estimates to the sampling costs. The measurement of the upper diameter (d6) is of less value than is generally suggested.

The PDF includes a summary in Finnish.

  • Kilkki, ORCID ID:E-mail:
article id 7620, category Article
Min-Sup Chung. (1981). Biochemical methods for determining population structure in Pinus sylvestris L. Acta Forestalia Fennica no. 173 article id 7620. https://doi.org/10.14214/aff.7620

Studies on Finnish Scots pine (Pinus sylvestris L.) plus tree clones by monoterpene and isozyme analyses was undertaken to further investigate mating system, population structure and pollination. Six allozyme systems (3 GOT, 1 GDH and 2 LAP) were properly analysed on the basis of segregation. Monoterpenes were analysed from needle material and segregation in high and low 3-carene content was found to depend on two alleles C and c. Thus, six allozyme systems and one monoterpene system were used as markers in this study.

It was shown that the northern clonal group maintains a much genetic variation as the central or southern clonal groups. The conditional probability of self-fertilization in about 20-year old clones estimated by the multilocus model was 14.1%, of which 8% originate from mating between trees that carry the same alleles to one of the maternal parent at some loci and 6% through self-fertilization.

There was no prominent difference in allele frequency of male gametes that pollinated the very early or very late flowering clones. The northern clonal group has higher a lower frequency of alleles GOT B2 and B3 respectively than of the southern clonal groups. The artificial plus tree selection, particularly in northern Finland, appears to favour heterozygous genotypes for the alleles that control 3-carene content n Scots pine.

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  • Chung, ORCID ID:E-mail:
article id 7590, category Article
Jukka Sarvas. (1977). Mathematical model for the physiological clock and growth. Acta Forestalia Fennica no. 156 article id 7590. https://doi.org/10.14214/aff.7590

In this paper a precise definition is given of the term physiological clock and the role of this clock in biological developmental and growth processes is mathematically studied. The heat sum method employed in the study of the annual cycle of development of forest trees has been used as the starting point. The mathematical principles of this method are analysed and it is shown that, on the same principles, a fairly general physiological clock can be constructed. Also, two growth models are presented in which this generalized physiological clock proves to play an important role.

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  • Sarvas, ORCID ID:E-mail:
article id 7572, category Article
Tauno Kallio, Pekka Tamminen. (1974). Decay of spruce (Picea abies (L.) Karst.) in the Åland Islands. Acta Forestalia Fennica no. 138 article id 7572. https://doi.org/10.14214/aff.7572

In 1972, all Norway spruce (Picea abies (L.) H. Karst.) trees of a minimum 7 cm diameter at breast height growing in the sample plots of the Sixth National Forest Inventory were examined on the main island of Aland, Finland. The soundness of standing trees was estimated by means of external characteristics and increment borer chips. The trees were then felled and measured. They were cut into lengths, and the type and extent of decay were studied.

30% of the trees examined was affected by butt rot, ca. 3% by wound decay. A comparison of the results with those of the Sixth National Forest Inventory justifies the estimate that in Aland 23% of spruce trees exceeding 7 cm in diameter at 1.3 m had butt rot.

The proportion of decayed trees in the cubic volume was 31%. Decayed wood material accounted for 5% of the volume including bark. Butt rot increased towards the mature stands. The reduction in the number of timber trees due to decay was 14.5%, in their volume 21.5%, and in the volume of sulphite pulpwood 12%. The share of sulphate pulpwood increased from 1 to 10%. The total reduction in usable wood was 6.3%. The stumpage price of the trees fell by 10.3%. As the degree of decay increased the increment percentage of the trees decreased. The most common cause of butt rot was Fomes annosus (Heterobasidion annosum) found in 46% of the number of decayed trees. Armillaria mellea was found in 16%. Bacteria were found in 50% of the decayed trees.

The PDF includes a summary in Finnish.

  • Kallio, ORCID ID:E-mail:
  • Tamminen, ORCID ID:E-mail:
article id 4644, category Article
Vilho Antero Kolehmainen. (1955). Havaintoja kulotuksen merkityksestä metsiemme uudistamisessa. Silva Fennica no. 85 article id 4644. https://doi.org/10.14214/sf.a9106
English title: Effect of prescribed burning in the forest regeneration.

Prescribed burning has reported to avail forest regeneration, for instance, by releasing nutrients for the use of seedlings, changing the pH of the soil and decreasing competition of ground vegetation. The aim of the study was to find out if the effects could be verified. Sample plots were measured in the experimental area of Tuomarniemi, in Central Finland, both in previously burned and untreated seedling stands and young forests. The main species in the sample plots was Scots pine (Pinus sylvestris L.).

According to the results, prescribed burning prepares the soil for regeneration. Germination percentage of the seeds is higher on the burned soil. All the species, Scots pine, Norway spruce (Picea abies (L.) Karst.) and birch species (Betula sp.) grow faster. Prescribed burning increases the amount of birch seedlings by improving its regeneration compared to unburned sites. The seed trees survive burning better if they are tall and have short crown, and have thick bark. In general, prescribed burning improves regeneration in seed tree stands.

The article includes a summary in German.

  • Kolehmainen, ORCID ID:E-mail:
article id 4627, category Article
Paavo Jaakko Ollinmaa. (1952). Jalot lehtipuumme luontaisina ja viljeltyinä. Silva Fennica no. 77 article id 4627. https://doi.org/10.14214/sf.a9099
English title: Native and cultivated southern broadleaved tree species in Finland.

The aim of the study was to update knowledge of natural range of English oak (Quercus robur L.), European ash (Fraxinus exelsior L.), Norway maple (Acer platanoides L.), small-leaved lime (Tilia cordata Miller), wych elm (Ulmus glabra Mill.) and European white elm (Ulmus laevis Pall.) in Finland, and estimate how far north they could be grown as forest trees or as park trees. The study is based on literature and questionnaires sent to cities and towns, District Forestry Boards, districts of Forest Service, Forestry Management Associations and railway stations.

The northern borders in the natural range of the species succeed one another from south to north as follows: English oak, European ash, Norway maple, wych elm, and small-leaved lime. Occurrence of European white elm is sporadic. The English oak forms forests in the southernmost Finland, while the other species grow only as small stands, groups or solitary trees. According to experiences of planted stands or trees, the northern limits of the species succeed one another from south to north as follows: European ash, English oak, Norway maple, European white elm, wych elm and small-leaved lime. All the species are grown in parks fairly generally up to the district of Kuopio-Vaasa (63 °). The northern limits where the species can be grown as park trees reach considerably further north in the western part of the country than in the east.

The article includes a summary in English.

  • Ollinmaa, ORCID ID:E-mail:
article id 4627, category Article
Paavo Jaakko Ollinmaa. (1952). Jalot lehtipuumme luontaisina ja viljeltyinä. Silva Fennica no. 77 article id 4627. https://doi.org/10.14214/sf.a9099
English title: Native and cultivated southern broadleaved tree species in Finland.

The aim of the study was to update knowledge of natural range of English oak (Quercus robur L.), European ash (Fraxinus exelsior L.), Norway maple (Acer platanoides L.), small-leaved lime (Tilia cordata Miller), wych elm (Ulmus glabra Mill.) and European white elm (Ulmus laevis Pall.) in Finland, and estimate how far north they could be grown as forest trees or as park trees. The study is based on literature and questionnaires sent to cities and towns, District Forestry Boards, districts of Forest Service, Forestry Management Associations and railway stations.

The northern borders in the natural range of the species succeed one another from south to north as follows: English oak, European ash, Norway maple, wych elm, and small-leaved lime. Occurrence of European white elm is sporadic. The English oak forms forests in the southernmost Finland, while the other species grow only as small stands, groups or solitary trees. According to experiences of planted stands or trees, the northern limits of the species succeed one another from south to north as follows: European ash, English oak, Norway maple, European white elm, wych elm and small-leaved lime. All the species are grown in parks fairly generally up to the district of Kuopio-Vaasa (63 °). The northern limits where the species can be grown as park trees reach considerably further north in the western part of the country than in the east.

The article includes a summary in English.

  • Ollinmaa, ORCID ID:E-mail:

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