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Fig. 1. The locations of the 34 tree breeding field trials of silver birch in southern and central Finland.

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Fig. 2. Latitudinal seed transfer distances of the seed orchard and unimproved progenies tested in the 34 silver birch field trials in southern and central Finland.

Table 1. The 19 silver birch seed orchards represented in the 34 common-garden trials. Origin means the geographical origin of seed orchard clones in Finland.
Orchard Alias Origin Type Clones Area ha Establishment year,
production years
Trials Offspring
288 South-Central Seedling 120 0.14 1972, 1973–1978 3 1491
336 Central Clonal 115 0.14 1977, 1979–1987 19 23 715
362 JR-1 Southern Clonal 2 0.03 1979, 1982–1996 26 1493
363 JR-1 Southern Clonal 2 0.04 1985, 1986–1993 26 1493
364 Southern Seedling 50 0.08 1985, 1987–1991 10 3334
373 Central Clonal 47 0.15 1987, 1989–1996 15 13 714
377 JR-1 Southern Clonal 2 0.05 1989, 1992–1996 26 1493
378 Southern Clonal 33 0.10 1989, 1992–1997 7 685
379 Central Clonal 32 0.20 1989, 1990–1993 4 578
381 Southern Clonal 35 0.10 1990, 1991–1998 4 566
383 JR-2 Southern Clonal 2 0.06 1990, 1991–1998 19 1491
384 JR-1 Southern Clonal 2 0.06 1990, 1991–1998 26 1493
387 Southern Clonal 41 0.05 1991, 1993–2003 3 348
390 Central-North Clonal 50 0.08 1992, 1994–2000 2 291
393 Central Clonal 69 0.07 1992, 1995–2002 2 2067
422 Central Clonal 78 0.14 1998, 2001–2004 8 516
424 Central Clonal 49 0.05 1992, 1995–2003 2 1456
430 Central Clonal 48 0.15 1998, 2001–2009 8 525
431 Southern Clonal 35 0.11 1999, 2003–2009 3 353
Table 2. Realised genetic gains from 19 silver birch seed orchards in tree height (H), stem diameter (DBH), stem volume (VOL), stem slenderness (HD) and number of ramicorn branches (NRAM) and forks (NFOR) as percentages of the unimproved mean. The standard errors of the gains are given within parentheses. The bold values were found significant at p < 0.05 of Dunnett’s test. In the seed orchard column, “JR-1” refers to the four seed orchards Nos. 362, 363, 377, and 384, while “JR-2” refers to seed orchard No. 383.
Orchard H DBH VOL HD NRAM NFOR
JR-1 3.9 (0.8) 3.2 (1.1) 8.9 (2.7) 0.2 (0.9) –6.1 (4.1) –42.7 (9.5)
JR-2 11.1 (0.7) 4.0 (0.9) 18.9 (2.4) 7.3 (0.7) –7.5 (4.1) –57.6 (10.4)
288 3.5 (1.7) 2.7 (1.9) 8.3 (5.1) 1.2 (1.4) –9.5 (7.4) –0.7 (16.9)
336 4.1 (0.7) 2.2 (0.8) 7.7 (2.1) 1.8 (0.6) –6.0 (2.2) –3.6 (4.1)
364 3.8 (1.0) 3.5 (1.3) 11.7 (3.2) 0.7 (0.9) –9.3 (3.8) –10.4 (8.3)
373 6.8 (0.8) 5.3 (0.9) 17.4 (2.3) 1.6 (0.6) –9.2 (2.3) –11.0 (3.9)
378 4.2 (1.1) 0.6 (1.4) 6.1 (3.5) 4.2 (1.1) –14.4 (7.7) –4.7 (11.4)
379 3.6 (1.3) 3.3 (1.6) 11.2 (4.3) –0.4 (1.3) 1.6 (8.8) –3.8 (14.6)
381 6.7 (1.3) 3.8 (1.6) 13.6 (4.3) 3.3 (1.3) 11.6 (8.4) 4.3 (13.8)
387 7.4 (1.6) 6.1 (1.9) 18.2 (4.9) 1.1 (1.4) 4.7 (7.9) –41.1 (18.2)
390 1.9 (2.2) 1.7 (2.1) 1.0 (7.3) 4.5 (2.4)
393 2.8 (2.0) 3.4 (1.7) 6.3 (6.2) 1.3 (1.9)
422 5.1 (1.1) 5.3 (1.3) 14.1 (3.6) 1.0 (1.1) –6.4 (7.4) –34.1 (17.0)
424 1.6 (2.0) 2.5 (1.7) 5.4 (6.2) 0.5 (1.9)
430 7.0 (1.1) 9.4 (1.3) 23.9 (3.6) –1.0 (1.1) –26.7 (8.4) –9.1 (14.7)
431 12.8 (1.6) 9.2 (1.9) 31.1 (4.8) 3.5 (1.4) –10.9 (8.4) –17.6 (15.5)
Average 5.4 4.1 12.7 1.9 –6.8 –16.2
Table 3. D-values from 19 silver birch seed orchards for tree height (H), stem diameter (DBH), stem volume (VOL), overall quality (OQns), branch quality (BQns), stem straightness (SSns), number of ramicorns (NRAMns) and number of forks (NFORns). The values indicate the probability of a randomly chosen seed orchard tree having a greater trait value than a random unimproved tree. In the seed orchard column, “JR-1” refers to the four seed orchards Nos. 362, 363, 377, and 384, while “JR-2” refers to seed orchard No. 383.
Orchard H DBH VOL HD OQns BQns SSns NRAMns NFORns
JR-1 0.60 0.55 0.56 0.50 0.58 0.62 0.52 0.48 0.34
JR-2 0.77 0.56 0.63 0.63 0.71 0.73 0.63 0.48 0.27
288 0.59 0.54 0.56 0.52 0.57 0.56 0.53 0.47 0.50
336 0.61 0.53 0.55 0.53 0.57 0.57 0.52 0.48 0.49
364 0.60 0.55 0.58 0.51 0.53 0.53 0.54 0.47 0.47
373 0.68 0.58 0.62 0.53 0.59 0.61 0.54 0.47 0.47
378 0.61 0.51 0.54 0.57 0.60 0.59 0.56 0.45 0.49
379 0.60 0.55 0.58 0.49 0.58 0.59 0.56 0.50 0.49
381 0.68 0.56 0.59 0.56 0.57 0.62 0.52 0.53 0.51
387 0.69 0.59 0.62 0.52 0.52 0.51 0.35
390 0.55 0.53 0.51 0.58 0.61 0.57 0.52
393 0.58 0.55 0.54 0.52 0.56 0.58 0.54
422 0.64 0.58 0.60 0.52 0.60 0.63 0.59 0.48 0.38
424 0.54 0.54 0.54 0.51 0.58 0.56 0.53
430 0.68 0.64 0.66 0.48 0.62 0.56 0.46 0.41 0.47
431 0.81 0.64 0.71 0.56 0.53 0.47 0.44
Average 0.64 0.56 0.59 0.53 0.59 0.59 0.54 0.48 0.44
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Fig. 3. D-values of the silver birch seed orchards for stem volume, branch quality stem straightness, number of ramicorns, and number of forks. The seed orchards are arranged in the ascending order of the average stem volume and branch quality D-values, reflecting the overall value of the orchard.

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Fig. 4. The relationship between the establishment year of the silver birch seed orchard and the D-values for stem volume (VOL), branch quality (BQns), and the number of forks (NFORns).

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Fig. 5. The relationship between the number of clones in the silver birch seed orchards and the D-values for stem volume (VOL), branch quality (BQns), and the number of forks (NFORns).