Full text of this article is only available in PDF format.

Timo Kuuluvainen (email), Kari Leinonen, Markku Nygren, Antti Penttinen

Statistical opportunities for comparing stand structural heterogeneity in managed and primeval forests: an example from boreal spruce forest in southern Finland.

Kuuluvainen T., Leinonen K., Nygren M., Penttinen A. (1996). Statistical opportunities for comparing stand structural heterogeneity in managed and primeval forests: an example from boreal spruce forest in southern Finland. Silva Fennica vol. 30 no. 2–3 article id 5598. https://doi.org/10.14214/sf.a9243


The horizontal and vertical stand structure of living trees was examined in a managed and in a primeval Norway spruce-dominated forest in Southern Finland. Tree size distributions (DBHs, tree height) were compared using frequency histograms. The vertical distribution of tree heights was illustrated as tree height plots and quantified as the tree height diversity (THD) using the Shannon-Weaver formula. The horizontal spatial pattern of trees was described with stem maps and quantified with Ripley's K-function. The spatial autocorrelation of tree sizes was examined with semivariogram analysis. In the managed forest the DBH and height distributions of trees were bimodal, indicating a two-layered vertical structure with a single dominant tree layer and abundant regeneration in the understory. The primeval forest had a much higher total number of trees which were rather evenly distributed in different diameter and tree height classes. The K-function summaries for trees taller than 15 m indicated that the primeval stand was close to complete random pattern. The managed stand was regular at small distances (up to 4 m). The semivariograms of tree sizes (DBH tree height) showed that the managed forest had a clear spatial dependence in tree sizes up to inter-tree distances of about 12 meters. In contrast, the primeval spruce forest had a variance peak at very short inter-tree distances (< 1 m) and only weak spatial autocorrelation at short inter-tree distances (1–5 m). Excluding the understory trees (h < 15 m) from the analysis drastically changed the spatial structure of the forest as revealed by semivariograms. ln general, the structure of the primeval forest was both horizontally and vertically more variable and heterogeneous compared to the managed forest. The applicability of the used methods in describing fine-scale forest structure i discussed.

boreal forests; Norway spruce; biodiversity; Picea abies; stand structure; semivariance; spatial analysis; K-function; structural variation

Published in 1996

Views 1137

Available at https://doi.org/10.14214/sf.a9243 | Download PDF

Creative Commons License

Click this link to register to Silva Fennica.
Log in
If you are a registered user, log in to save your selected articles for later access.
Contents alert
Sign up to receive alerts of new content

Your selected articles
Your search results
Kuuluvainen T., Leinonen K. et al. (1996) Statistical opportunities for comparing stand st.. Silva Fennica vol. 30 no. 2–3 article id 5598
Leinonen K., Rita H. (1995) Interaction of prechilling, temperature, osmotic.. Silva Fennica vol. 29 no. 2 article id 5549
Leinonen K., Leikola M. et al. (1989) Natural regeneration of Norway spruce in Pirkka-.. Silva Fennica vol. 0 no. 209 article id 7656
Chantal M. d., Eskola L. et al. (2003) Early establishment of Pinus sylvestris and Pice.. Silva Fennica vol. 37 no. 1 article id 509