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Risto Heikkilä (email), Timo Mikkonen

Effects of density of young Scots pine (Pinus sylvestris) stand on moose (Alces alces) browsing

Heikkilä R., Mikkonen T. (1992). Effects of density of young Scots pine (Pinus sylvestris) stand on moose (Alces alces) browsing. Acta Forestalia Fennica no. 231 article id 7677. https://doi.org/10.14214/aff.7677

Abstract

The study was carried out at Padasjoki, Southern Finland, where moose (Alces alces L.) density on the winter range had been over 1.5 individuals/km2. Moose browsing intensity, expressed in terms of number of twigs eaten and biomass used, increased with stand density (biomass). Total biomass consumption (dry weight) per sample plot and per sapling. The number of bites increased, but the percentage biomass removed did not differ when stand density increased. A relatively large bite size was observed on the plots of low stand density. The quantity of food, which on average was of relatively low quality, was obviously important due to the benefit gained through reducing the search time.

The nutritive value of the browse, expressed in terms of chemical compounds indicating low food digestibility, was lower in the dense than in the sparse Scots pine stand. However, the amount of crude protein and arginine were relatively high in the dense stand. We concluded that shading affected the nutritional status of saplings on high density plots.

Although the biomass removed by moose per sapling was high for low density plots, the remaining biomass was larger than that on the high-density plots owing to the relatively large twig biomass of saplings. The number of saplings per hectare without main stem breakage increased significantly as stand density increased.

The PDF includes a summary in Finnish.

Keywords
Pinus sylvestris; Scots pine; Alces alces; stand density; feeding behaviour; moose browsing; plant phenology

Published in 1992

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Available at https://doi.org/10.14214/aff.7677 | Download PDF

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Kuuluvainen T., Pukkala T. (1989) Effect of Scots pine seed trees on the density o.. Silva Fennica vol. 23 no. 2 article id 5379
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Kellomäki S., Hänninen H. et al. (1987) A tentative model for describing the effects of .. Silva Fennica vol. 21 no. 1 article id 5302
Velling P., Nepveu G. (1986) Variation of wood quality and yield in a Finnish.. Silva Fennica vol. 20 no. 3 article id 5275
Löyttyniemi K., (1985) On repeated browsing of Scots pine saplings by m.. Silva Fennica vol. 19 no. 4 article id 5252
Kellomäki S., (1984) Observations on the influence of stand density o.. Silva Fennica vol. 18 no. 2 article id 5207
Heliövaara K., Annila E. et al. (1983) Effect of nitrogen fertilization and insecticide.. Silva Fennica vol. 17 no. 4 article id 5197
Rousi M., (1983) Susceptibility of pine to mammalian herbivores i.. Silva Fennica vol. 17 no. 4 article id 5195
Kellomäki S., (1983) Strength of Scots pine branches Silva Fennica vol. 17 no. 2 article id 5185
Mälkönen E., Aro-Heinilä V. et al. (1982) Effect of fertilization and irrigation on the gr.. Silva Fennica vol. 16 no. 1 article id 5157
Ollinmaa P. J., (1981) Physical properties of wood growing on drained s.. Silva Fennica vol. 15 no. 3 article id 5128
Kellomäki S., (1981) Effect of the within-stand light conditions on t.. Silva Fennica vol. 15 no. 2 article id 5111
Kellomäki S., Kanninen M. (1980) Eco-physiological studies on young Scots pine st.. Silva Fennica vol. 14 no. 4 article id 5092
Meriluoto J., (1980) Applicability of MCPA- and 2,4,5-T-herbicides in.. Silva Fennica vol. 14 no. 4 article id 5086
Kellomäki S., Hari P. et al. (1980) Eco-physiological studies on young Scots pine st.. Silva Fennica vol. 14 no. 3 article id 5079
Velling P., (1980) Variation in the density of wood of different Sc.. Silva Fennica vol. 14 no. 1 article id 5063
Kellomäki S., (1979) The effect of solar radiation and air temperatur.. Silva Fennica vol. 13 no. 4 article id 5048
Hari P., Kanninen M. et al. (1979) An automatic system for measurements of gas exch.. Silva Fennica vol. 13 no. 1 article id 5019
Kellomäki S., (1979) On geoclimatic variation in basic density of Sco.. Silva Fennica vol. 13 no. 1 article id 5015
Löyttyniemi K., Hiltunen R. (1978) Monoterpenes in Scots pine in relation to browsi.. Silva Fennica vol. 12 no. 2 article id 4992
Kärkkäinen M., (1977) Comparison of wood properties of Parana pine and.. Silva Fennica vol. 11 no. 1 article id 4958
Lehtonen I., Kellomäki S. et al. (1976) Nutrient cycle in a Scots pine stand. II. Amount.. Silva Fennica vol. 10 no. 4 article id 4952
Lehtonen I., Kellomäki S. et al. (1976) Nutrient cycle in a Scots pine stand I. Seasonal.. Silva Fennica vol. 10 no. 3 article id 4946
Kapustinskaité T., (1975) Ash content of peatland soils and stand growth i.. Silva Fennica vol. 9 no. 3 article id 4924
Päivänen J., (1973) The effect of thinning on the snow cover and soi.. Silva Fennica vol. 7 no. 2 article id 4881
Leikola M., (1971) Throughfall in a managed Scots pine stand in Sou.. Silva Fennica vol. 5 no. 2 article id 4842
Löyttyniemi K., (1968) Feeding of terminal shoots of Scots pine seedlin.. Silva Fennica vol. 2 no. 4 article id 4778
Pulliainen E., Loisa K. et al. (1968) Winter food of the moose (Alces alces) in easter.. Silva Fennica vol. 2 no. 4 article id 4775
Leikola M., (1967) Observations on wind conditions in a managed Sco.. Silva Fennica vol. 1 no. 3 article id 4754
Sainio P., (1955) Winter nutrition of elk Silva Fennica vol. no. 88 article id 4647
Heiskanen V., (1965) Relation between the development of the early ag.. Acta Forestalia Fennica vol. 80 no. 2 article id 7164
Heikurainen L., Seppälä K. (1965) Regionality in stand increment and its dependenc.. Acta Forestalia Fennica vol. 78 no. 4 article id 7157
Seppänen M., (1964) Distribution of rainfall in the Scots pine stand.. Acta Forestalia Fennica vol. 76 no. 8 article id 7148
Yli-Vakkuri P., (1961) Studies on the development of young sown pine st.. Acta Forestalia Fennica vol. 74 no. 3 article id 7130
Kallio K., (1960) The mensurational density of a stand in estimati.. Acta Forestalia Fennica vol. 71 no. 7 article id 7116
Jalava M., (1934) Influence of the position of a tree in the stand.. Acta Forestalia Fennica vol. 40 no. 9 article id 7286
Hari P., Ross J. et al. (1996) Production process of Scots pine Acta Forestalia Fennica vol. 0 no. 254 article id 7525
Luomajoki A., (1993) Climatic adaptation of Scots pine (Pinus sylvest.. Acta Forestalia Fennica vol. 0 no. 237 article id 7683
Heikkilä R., (1991) Moose browsing in a Scots pine plantation mixed .. Acta Forestalia Fennica vol. 0 no. 224 article id 7670
Lääperi A., (1990) Effect of winter feeding on moose damage to youn.. Acta Forestalia Fennica vol. 0 no. 212 article id 7659
Pohtila E., Pohjola T. (1983) The timing of foliage spraying during the growin.. Acta Forestalia Fennica vol. 0 no. 181 article id 7628
Nyyssönen A., Ojansuu R. (1982) Assessment of timber assortments, value and valu.. Acta Forestalia Fennica vol. 0 no. 179 article id 7626
Hari P., Kellomäki S. et al. (1982) Dynamics of early development of tree stand Acta Forestalia Fennica vol. 0 no. 177 article id 7624
Heikurainen Leo., (1980) Drainage condition and tree stands on peatlands .. Acta Forestalia Fennica vol. 0 no. 167 article id 7614
Nyyssönen A., Mielikäinen K. (1978) Estimation of stand increment Acta Forestalia Fennica vol. 0 no. 163 article id 7597
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Vacek S., Vacek Z. et al. (2016) Structure, regeneration and growth of Scots pine.. Silva Fennica vol. 50 no. 4 article id 1564
Ruuhola T., Nikula A. et al. (2016) Effects of bedrock and surficial deposit composi.. Silva Fennica vol. 50 no. 3 article id 1565
Moilanen M., Hytönen J. et al. (2015) Fertilization increased growth of Scots pine and.. Silva Fennica vol. 49 no. 3 article id 1301
Bergqvist G., Bergström R. et al. (2014) Recent browsing damage by moose on Scots pine, b.. Silva Fennica vol. 48 no. 1 article id 1077
Karlsson L., Mörling T. et al. (2013) Influence of silvicultural regimes on the volume.. Silva Fennica vol. 47 no. 4 article id 938
Terhonen E., Marco T. et al. (2011) The effect of latitude, season and needle-age on.. Silva Fennica vol. 45 no. 3 article id 104
Varis S., Pakkanen A. et al. (2009) The extent of south-north pollen transfer in Fin.. Silva Fennica vol. 43 no. 5 article id 168
Ikonen V.-P., Kellomäki S. et al. (2009) Sawn timber properties of Scots pine as affected.. Silva Fennica vol. 43 no. 3 article id 197
Gort J., Zubizarreta Gerendiain A. et al. (2009) Differences in fibre properties in Scots pine (P.. Silva Fennica vol. 43 no. 3 article id 193
Peltola H., Gort J. et al. (2009) Differences in growth and wood density traits in.. Silva Fennica vol. 43 no. 3 article id 192
Nikula A., Hallikainen V. et al. (2008) Modelling the factors predisposing Scots pine to.. Silva Fennica vol. 42 no. 4 article id 235
Saksa T., Miina J. (2007) Cleaning methods in planted Scots pine stands in.. Silva Fennica vol. 41 no. 4 article id 274
Wennström U., Bergsten U. et al. (2007) Seedling establishment and growth after direct s.. Silva Fennica vol. 41 no. 2 article id 298
Siipilehto J., (2006) Height distributions of Scots pine sapling stand.. Silva Fennica vol. 40 no. 3 article id 331
Salminen H., Jalkanen R. (2006) Modelling variation of needle density of Scots p.. Silva Fennica vol. 40 no. 2 article id 337
Sarkkola S., Hökkä H. et al. (2004) Natural development of stand structure in peatla.. Silva Fennica vol. 38 no. 4 article id 408
Kojola S., Penttilä T. et al. (2004) Impacts of different thinning regimes on the yie.. Silva Fennica vol. 38 no. 4 article id 407
Warensjö M., Rune G. (2004) Stem straightness and compression wood in a 22-y.. Silva Fennica vol. 38 no. 2 article id 424
Pukkala T., Miina J. et al. (2002) Thinning response and thinning bias in a young S.. Silva Fennica vol. 36 no. 4 article id 524
Jansons Ā., Robalte L. et al. (2016) Long-term effect of whole tree biomass harvestin.. Silva Fennica vol. 50 no. 5 article id 1661