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                            Herbaceous ground vegetation in artificially-created gaps was studied in  a managed beech (Fagus sylvatica L.) forest over a period of eight  years in Northern Hungary, Central Europe. These gaps were being used as  an alternative to the regular shelterwood system to create uneven-aged  stands. The effects of gap size (15 and 40 m diameter) and canopy  openness on herbaceous species colonization and persistence were  assessed in a systematic grid of 5   5 m. Overall, herbaceous cover was  low before gap creation, increased soon afterwards, and continued to  rise over time. The number of herb species increased in the gaps and, to  a lesser extent, in adjacent areas under the remaining tree canopy.  Colonization of gaps was rapid and there was substantial turnover of  species i.e. various species disappeared from the gaps over time whilst  others colonized. Species with both long-term persistent seed banks and  long distance dispersal abilities were the most successful types  colonizing gaps. Six species occurred preferentially in large gaps,  while only one species was found to prefer small gaps. Species present  before gap creation survived in both gap sizes. Smaller gaps with a  diameter of half the height of canopy trees also tended to remain free  of common weed species, whereas large cover of Rubus fruticosus L. and  Calamagrostis epigejos (L.) Roth could hamper natural regeneration in  larger gaps. For the successful regeneration of beech we recommend the  use of small gaps complemented by few large gaps.
                        
                
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                            Kelemen,
                            Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary
                                                        E-mail:
                                                            kk@nn.hu
                                                                                
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                            Mihók,
                            Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary
                                                        E-mail:
                                                            bm@nn.hu
                                                                                
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                            Gálhidy,
                            Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary
                                                        E-mail:
                                                            lg@nn.hu
                                                                                
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                            Standovár,
                            Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary
                                                        E-mail:
                                                            standy@ludens.elte.hu
                                                                                        