Silva Fennica vol. 58 no. 3 | 2024

• Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, Department of Forestry and Wildlife Management, Campus Evenstad, Postboks 400, 2418 Elverum, Norway E-mail: sjoliehannek3@gmail.com

Mixed forests are known for their ability to provide a wide range of ecosystem services. Such forests have higher biodiversity compared to monocultures, are resilient against disturbances and may mitigate the effects of climate change. Despite well-known benefits, there is still little information on how these forests should be established and managed. The aim of this study was to describe the early growth dynamics of current boreal young mixed stands of planted Norway spruces (Picea abies (L.) Karst.) and naturally regenerated birches (Betula spp.). We collected data from 9  stands planted for spruce 8–14 years ago in Southern and Central Finland. Stem analysis was conducted to 144 spruces and to 144 birches to determine previous growth. We modelled the height and diameter development of individual trees in relation to tree age at stump height using non-linear mixed Chapman-Richards model. There were no significant differences between spruce and seed-origin birch in diameter growth at stump height, but the initial height increments of natural birches were larger than those of planted spruces. However, planted spruces were able to keep up with the height development of birches, if spruces received a head start over naturally regenerated seed-origin birch for two growing seasons. Thus, naturally regenerated birch admixture can be utilized to establish single-storied spruce-birch mixtures, and the admixture should be retained during the early cleaning of planted spruce stands.

• Männistö, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0009-0009-0213-1972 E-mail: lauri.mannisto@luke.fi
• Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
• Huuskonen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi

Disturbances caused by the European spruce bark beetle (SBB; Ips typographus L.) on Norway spruce (Picea abies (L.) H. Karst.), have increased immensely across Central and Northern Europe, and are expected to increase further as a result of climate change. While this trend has been noted in Finland, so far limited research has been published. To support proper SBB risk management in Finland, we compared stand properties between salvage loggings due to SBB damage during 2012–2020 (4691 cases) and spruce stands free of SBB damage. Also, we explored the role of landscape attributes as drivers of SBB damage. We considered the forest stand attributes of site fertility class, stand development class, soil type, stand mean diameter at breast height and mean stand age. Considered forest landscape attributes were the distance from SBB-damaged stands to the closest clear-cut, to previous-year SBB-damaged stands and to the previous-year wind-damaged stand. We used nationwide forest logging and forest stock data, and analysed forest stand attributes using chi-squared and Mann-Whitney U tests and landscape attributes using generalised linear mixed models. Based on our findings, the SBB didn’t damage stands randomly, but prevailed in mature stands (high age and high mean diameter at breast height), in herb-rich heath forest site types and in semi-coarse or coarse heath forest soil soils. We found correlation between the landscape variables and the number of salvage loggings, with a higher number of loggings due to SBB damage close to clear-cuts. Our results help to find risk areas of SBB damage.

• Pulgarin Diaz, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-0554-8254 E-mail: alexander.pulgarin.diaz@uef.fi
• Melin, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80101 Joensuu, Finland https://orcid.org/0000-0001-7290-9203 E-mail: markus.melin@luke.fi
• Ylioja, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-0079 Helsinki, Finland https://orcid.org/0000-0002-8840-7504 E-mail: tiina.ylioja@luke.fi
• Lyytikäinen-Saarenmaa, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-1884-3084 E-mail: paivi.lyytikainen-saarenmaa@ef.fi
• Peltola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
• Tikkanen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-3875-2772 E-mail: olli-pekka.tikkanen@uef.fi

Forest fires pose a significant threat to forest carbon storage and sinks, yet they also play a crucial role in the natural dynamics of boreal forests. Accurate quantification of biomass changes resulting from forest fires is essential for damage assessment and controlled burning evaluation. This study utilized terrestrial laser scanning (TLS) to quantify changes in ground vegetation resulting from low-intensity surface fires. TLS data were collected before and after controlled burnings at eight one-hectare test sites in Scots pine (Pinus sylvestris L.) dominated boreal forests in Finland. A surface differencing-based method was developed to identify areas exposed to fire. Validation, based on visual interpretation of 1 × 1 m surface patches (n = 320), showed a recall, precision, and F1-score of 0.9 for the accuracy of identifying burned surfaces. The developed method allowed the assessment of the magnitude of fire-induced vegetation changes within the test sites. The proportions of burned 1 × 1 m areas within the test sites varied between 51–96%. Total volumetric change in ground vegetation was on average –1200 m³ ha^-1, with burning reducing the vegetation volume by 1700 m³ ha^-1 and vegetation growth increasing it by 500 m³ ha^-1. Substantial variations in the volumetric changes within and between the test sites were detected, highlighting the complex dynamics of surface fires, and emphasizing the importance of having observations from multiple sites. This study demonstrates that bitemporal TLS measurements provide a robust means for characterizing fire-induced changes, facilitating the assessment of the impact of surface fires on forest ecosystems.

• Tienaho, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0002-6574-5797 E-mail: noora.tienaho@uef.fi
• Saarinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0003-2730-8892 E-mail: ninni.saarinen@uef.fi
• Yrttimaa, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0003-2648-523X E-mail: tuomas.yrttimaa@uef.fi
• Kankare, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0001-6038-1579 E-mail: ville.kankare@uef.fi
• Vastaranta, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0001-6552-9122 E-mail: mikko.vastaranta@uef.fi

Uncrewed aerial vehicles (UAV) have great potential for use in forest inventories, but in practice they can be expensive for relatively small inventory areas as a large number of field measurements are needed for model construction. One proposed solution is to transfer previously constructed models to a new inventory area and to calibrate these with a small number of local field measurements. Our objective was to compare calibration of general models and the construction of new models to determine the best approach for UAV-based forest inventories. Our material included field measurements and UAV-based laser scanning data, from which individual trees were automatically identified. A general mixed-effects model for diameter at breast height (DBH) had been formulated earlier based on data from a geographically wider area. It was calibrated to the study area with field measurements from 2–10 randomly selected calibration trees. The calibrated diameters were used to calculate the diameter of a basal area median tree (DGM), tree volumes, and the volume of all trees at plot-level. Next, new DBH-models were formulated based on the 2–10 randomly selected trees and calibrated with plot-level random effects estimated during model construction. Finally, plot-specific height-diameter regression models were formulated by randomly selecting 10 trees from each plot. Calibration reduced the prediction errors of all variables. An increase in the number of calibration trees decreased error rates by 1–6% depending on the variable. Calibrated predictions from the general mixed-effects model were similar to the separately formulated mixed-effects models and plot-specific regression models.

• Jääskeläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0009-0004-4127-7863 E-mail: johanna.jaaskelainen@uef.fi
• Korhonen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9352-0114 E-mail: lauri.korhonen@uef.fi
• Kukkonen, Natural Resources Institute Finland, Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-4206-1680 E-mail: mikko.kukkonen@luke.fi
• Packalen, Natural Resources Institute Finland, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-1804-0011 E-mail: petteri.packalen@luke.fi
• Maltamo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9904-3371 E-mail: matti.maltamo@uef.fi

Reproduction in masting species is characterised by long intervals between good cone and seed production years, and only sparse reproduction between mast years. The physiological mechanisms behind masting, and how these are linked to internal resource status and external weather factors, is still a subject of scientific exploration and debate, as is the effect of climate change on masting. This study investigates cone production in one operational seed orchard in Sweden which was established with two different spacings and has since been subject to three tree thinning experiments. The spacings before thinning varied between 800 and 400 stems ha^–1, and then thinning reduced the stand density in all trials to half, i.e. between 400 and 200 stems ha^–1. In all three experiments cone production per tree was equal in un-thinned and thinned treatments, both in mast years and in non-mast years. Thus, the cone production per unit area was twice as high in the un-thinned areas. The conclusion from these experiments is that the establishment of Picea abies (L.) H. Karst. seed orchards with wide tree spacing is both a misuse of good orchard locations and bad economics.

• Almqvist, Skogforsk, Uppsala Science Park, SE-75183 Uppsala, Sweden https://orcid.org/0000-0001-5739-4854 E-mail: curt.almqvist@skogforsk.se

We evaluated the consistency of video, ordinary photo, and panoramic photo surveys in measuring the attractiveness (recreational use, scenic values etc.) of forest stands managed with varying intensities. We also evaluated possible effects on the results caused by the personal background of citizen respondents and how the respondents experienced the evaluation events. Our experimental sites were in mature Scots pine (Pinus sylvestris L.) forests in eastern Finland and included two replicate sites which were unharvested (control, basal area 26 m² ha^–1), a selective cutting site (basal area 18 m² ha^–1), small openings sites (gap cut) with 5 and 20% retained trees, respectively, and one site which was clear cut with 3% retained trees. In our study, 71 volunteer forestry students evaluated the attractiveness of these sites from an ordinary photo, a panoramic photo, and a video, with a 0–10 scale. Based on this study, the unharvested forest was the most attractive and clear cutting was the least attractive, regardless of the evaluation method. This result was in line with a previous study using on-site evaluations of the same sites. The differences of respondents considering in how easy they felt to assess the attractiveness of the environment as a whole and in using different visualisation methods affected the result, unlike background variables of the respondents. The results of forest attractiveness were consistent between panoramic and ordinary photos, and the attractiveness scoring was slightly higher for them than for the video. We conclude that all the compared visualisation methods seem to be suitable for assessment of the attractiveness of forest views.

• Silvennoinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9095-7986 E-mail: harri.silvennoinen@uef.fi
• Pikkarainen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0001-5301-3639 E-mail: laura.pikkarainen@uef.fi
• Nakola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heini.nakola@gmail.com
• Koivula, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-6415-4904 E-mail: matti.koivula@luke.fi
• Tyrväinen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-5144-7150 E-mail: liisa.tyrvainen@luke.fi
• Tikkanen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9693-8209 E-mail: jukka.tikkanen@uef.fi
• Chambers, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-0586-3142 E-mail: philip.chambers@uef.fi
• Peltola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi

The EU’s influence on national forest policies is growing, and the implementation of forest-related policies proposed by the Commission will affect the practice of forestry in Europe. For instance, the Nature Restoration Law sets concrete areal goals for restoring forest ecosystems and for conservation, the Deforestation Regulation requires meticulous tracking of wood’s origin, and the renewed Renewable Energy Directive (RED III) sets new criteria to sustainable forest biomass procurement. So far there have been no studies that have looked into the impacts from the economic and operational point of view. In this study, structural systems analysis was first performed to discover the relevant variables (and their functioning) associated with the roundwood harvesting operations and the operating environment. A scenario approach was then applied to capture the potential levels of implementation of the EU’s forest-related policies. Finally, using different scenarios (low-, moderate- and high-impact) and a systems analysis framework, the impact of alternative levels of implementation was quantified in terms of harvesting costs, measured in € m^–3. The results indicate that with the low- and moderate-impact scenarios the harvesting costs would increase by less than 10% from the current levels in three different regions in Finland. Such an increase (less than 10%) could be tolerated over a period of a few years, but a sudden increase is likely to lead to challenges to the running of businesses. With the high-impact scenario the harvesting costs would increase by between 15% and 18%, depending on the region. This magnitude of increase (of approximately a sixth) corresponds to a severe change in the roundwood harvesting operations and operating environment.

• Ahtikoski, Natural Resources Institute Finland (Luke), Tekniikankatu 1, FI-33720 Tampere, Finland https://orcid.org/0000-0003-1658-3813 E-mail: anssi.ahtikoski@luke.fi
• Väätäinen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-6886-0432 E-mail: kari.vaatainen@luke.fi
• Anttila, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi
• Laitila, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: juha.laitila@luke.fi
• Mutanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-0533-9356 E-mail: antti.mutanen@luke.fi
• Lindblad, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0009-0003-6766-6587 E-mail: jari.lindblad@luke.fi
• Sikanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: lauri.sikanen@luke.fi
• Routa, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-7225-1798 E-mail: johanna.routa@luke.fi

In boreal forests fires often ignite and spread within the dominant moss and lichen cover of the ground layer vegetation, which thus greatly influences fire hazard. We used an experimental set-up in greenhouse conditions to study the differences in how (1) fuel moisture and (2) wind velocity influence the ignition probability and fuel consumption among four common circumboreal ground vegetation fuels, Pleurozium schreberi (Willd. ex Brid.) Mitt., Hylocomium splendens Schimp., Dicranum spp. and Cladonia rangiferina (L.) F. H. Wigg. Our results show that the reindeer lichen C. rangiferina was clearly the most flammable species, with high ignition probability even at high moisture contents and low wind velocities. Of the mosses, Dicranum was the least flammable, with low ignition probability and mass loss at low wind velocities regardless of moisture content. P. schreberi and H. splendens behaved somewhat similarly with wind velocities quickly increasing the initially low ignition probability and mass loss observed in the absence of wind. However, especially for mass loss, among-species differences tended to disappear with stronger winds. The observed differences can be explained by the different structures and growth forms of the studied species and open a potential avenue for improving forest fire risk predictions.

• Lindberg, Häme University of Applied Sciences, School of Bioeconomy, Visamäentie 35 A, P.O. Box 230, FI-13100 Hämeenlinna, Finland https://orcid.org/0000-0001-9045-2673 E-mail: henrik.lindberg@hamk.fi
• Aakala, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-0160-6410 E-mail: tuomas.aakala@uef.fi
• Vanha-Majamaa, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: ilkka.vanha-majamaa@luke.fi