Current issue: 58(4)
Draining transforms root systems of trees growing in peatlands towards the ones growing on mineral soil. However, even after efficient draining the root systems differ from the root systems of trees growing on mineral soil. This investigation concentrates on root systems of forests of similar mire types growing in similar draining conditions but having different tree species compositions. The peatland, situated in Pieksämäki in Southern Finland, was drained in 1937. Sample plots, measured in 1956, consisted of mixed forest of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies L. Karst.) and birch (Betula sp.) in different compositions, and were in natural condition.
The sedge pine bog studied in this investigation was shown to have larger total amount of roots and mycorrhiza than in previously studied dwarf shrub pine bogs. This reflects better growth conditions of the better site. The depth of root system was, however, similar. Root systems of birch were deeper than those of the coniferous tree species. Differences between Scots pine and Norway spruce were small. Corresponding differences between the species were found in the density and total number of mycorrhizas. The abundance of mycorrhizas in the roots of birch increased in deeper layers of peat, but decreased especially in spruce roots. In earlier studies the abundance of mycorrhizas decreased in the roots growing in deeper layers in pure Scots pine stands, but no such variation was seen in this study. The result suggest that the deep root system of birch may affect also the root systems of the coniferous trees. On the other hand, birch roots can have advantage over the coniferous trees.
The PDF includes a summary in German.
According to studies following the development of vegetation of drained peatlands, it seems that they have transformed to a relatively stable plant communities during the succession. In earlier studies it was assumed that after drainage a mire type would develop to a corresponding forest site. This investigation studies what kinds of plant communities are formed during succession of different mire types on peatlands drained for forestry in the southern half of Finland. Understorey vegetation was studied in 18 sample plots established by Forest Research Institute on drained peatlands. In addition, sample plots were studied on peatlands in natural state.
The results suggest that understorey vegetation on peatlands drained for forestry have developed into plant communities, the most advanced of which are the so-called dry plant communities. They represent transformed site types, which are the following: drained peatlands with upland herb-rich vegetation, drained peatlands with upland grass-herb vegetation, drained peatlands with upland Myrtillus site type vegetation, drained peatlands with upland Vaccinium site type vegetation, and drained peatlands with upland Calluna site type vegetation. Drained peatlands with upland Cladonia site type vegetation seem to be a temporary type caused by incomplete drainage. The transition between Myrtillus and Vaccinium dominated dry plant communities is not clear, but especially the pure Vaccinium vitis-ideae communities justify its place as an independent plant community. The dry drwarf shrub plant communities are also stable.
The PDF includes a summary in German.
Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.
Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.
The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.
The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.
The PDF includes a summary in German.
According to the theory of peatland types, particular peatland types, after sufficient drainage, change into certain forest types. It has been found, that the range of forest types in peatlands in different stages of draining is as large as on mineral soil; and comprise Cladina, Calluna, Vaccinium, Myrtillus Oxalis-myrtillus and grove types. Poor peatland types change into poor forest types, better for better types. However, a Swedish scientist Mellin suggested that after effective drainage especially oligotrophic bogs, when well drained, change usually into Myrtillus type.
The different conclusions are due to the fact that the same bog type may develop into different forest types according to the effectiveness and duration of the drainage. Greater the decay of the peat bog layer, the more exacting is the type of vegetation which appears. Bog types of classes V, IV and III (Finnish classification of site quality) change into a Myrtillus type, as do the poorer peatland types of class II. The types vary, however, in their economical drainage value. The fact that bogs which in their natural stage are clearly different in their site quality change after through drainage into the same forest type, is explained by the chemical quality of the peat. However, class I and the best types of class II bogs change into better forest types because they as eutropchic bogs are richer in nitrogen and lime. This difference persists despite of effective draining.
It has been shown that the development of eutrophic peatland types at the forest type stage also differs clearly from the development of oligotrophic peatland types. The Finnish classification of drainage value shows correctly the relative drainability when using normal spacing of ditches. The notes on forest types on mineral soil should, however, be replaced by corresponding notes on the transitive types between bog and forest types.
The PDF includes a summary in Swedish and English.
The aim of the investigation was to study natural regeneration of Norway spruce (Picea abies (L.) Karst.) in drained peatlands and frost injuries in seedlings, and to compare microclimates of the regeneration areas. The experiments included peatlands in Satakunta in Western Finland. Restocking of the areas with seedlings and their survival was followed in 1935-40 at sample plots that were mainly 1 are large.
Susceptibility to freezing was shown to be dependent on the stage of development of the shoots. Shoots that have just begun to grow contain little water, and withstand better freezing temperatures than shoots in later stages of growth. Damages to the seedlings were observed when the temperatures decreased to -2.8–-4.3 °C. The most severe damage to a seedling was caused by the death of the leading shoot by spring frost.
Norway spruce regenerates easily on moist peatlands, but peatlands with dry surface tend to have little or no seedlings. The species regenerated better in marshy sites than correspondingly fertile mineral soil sites. However, it needs shelter to avoid frost damage. On clear cut spruce swamp the undergrowth spruce seedlings that were left in the site got severe frost damage. If the site had birch (Betula sp.) coppice or undergrowth, spruce seedlings survived in their shelter depending on the height and density of the birch trees. To be effective, the protective forest should have relatively even crown cover. Young spruce seedlings could grow well even under relatively dense birch stand.
The PDF includes a summary in German.
The drained peatlands regenerate usually well, and artificial regeneration by sowing or planting has been rare. Field trials of Norway spruce (Picea abies (L.) H. Karst.) were established in northern Satakunta in Western Finland in three drained peatlands in 1934. Sowing trials of Norway spruce consisted of patch and broadcast sowed sample sites in treeless bogs and under protective forest. The seedlings of spruce were planted either under protective forest or in treeless peatland.
The results show that artificial regeneration of Norway spruce succeeds best under protective forest. The best tree species for upper storey is Betula sp. which grows fast and controls growth of ground vegetation. The peat is relatively decomposed on those peatlands that are suitable for spruce, and breaking of the surface of the peat is not recommended. In the sowing trials, breaking of the upper layer of the peat caused frost heaving, cracking of the dried surface and sticking of mud in the seedlings in the patch sown sample site. The shoot and root growth of seedlings of the broadcast sown site was better than seedlings of the patch sown site. The planted spruce seedlings seemed to be more susceptible for spring frost than the seedlings in the sown site. The plants of seed origin succeeded in general better than the planted seedlings.
The PDF includes a summary in German.
Only about 24,000 hectares of peatlands have been drained in the state lands by the 1921. The aim of this study was to define how much the growth of the trees in the drained peatland revives. Sample plots were measured in previously drained peatlands that had sufficient Scots pine (Pinus sylvestris L.) tree stand. A stem analysis was performed to one of the sample trees. The evenness of the stands was dependent on how evenly the peatlands had dried when the stand was regenerated. Thus, the sample stands were not always fully stocked. However, they had capacity to develop towards evenly structured forests as the peatlands continued to dry further. The diameter and height growth of the dried peatlands have corresponded the similar stands in mineral soil sites. In trees that have grown stunted in the peatlands, the diameter growth seems to increase faster than the height growth. The volume growth is slightly smaller than in the similar mineral soil sites due to less favorable stem form. After the draining, the roots of the trees continued to grow from the old branches of root, but start then to form new roots. When the ground water level drops, the root layer grows deeper.
The PDF includes a summary in German.
Draining of peatlands to improve forest growth started to increase in Finland in the beginning of 1900s. The aim of the study was to find out which kind of peatlands are suitable for draining. The peatlands examined in this study had been drained earlier in 1800s for other purposes, and the original peatland type was deduced afterwards. When the peatland is drained, its vegetation changes gradually towards that of mineral soil sites, depending on the original peatland type. The article includes detailed description of the vegetation on different drained peatland sites. Best represented in the study were different types of pine swamps, which change towards Calluna or Vaccinium forest site type depending on the original peatland type. The Sphangnum species and brushwood disappear gradually and Cladina sp. become common in some drained pine swamp types. Scots pine (Pinus sylvestris L.) regenerates well on most drained pine swamps, and also Betula sp may grow as dominant species. The richer pine swamp types develop to Vaccinium-myrtillus forest site type, which may grow also Norway spruce (Picea abies (L.) H. Karst). Drained treeless bogs change first towards pine swamps. However, trees regenerate poorly on these sites and the growth is low. Flark-bogs develop typically to treeless lichen heaths. Drained spruce swamps develop to forest with grass-herb vegetation or Myrtillus site type.
The PDF includes a summary in German.
The paper presents some preliminary results of a 10-year-old study the purpose of which is to determine the effect of simultaneous variations in the intensity of drainage and fertilization on the development of planted and natural seedlings on peatlands under various climatic conditions. The development of the Scots pine (Pinus sylvestris L.) seedlings appeared to be better the more intensive the degree of drainage and fertilization used. The increase in the temperature sum had a positive effect on the development of pine seedlings and decreased the mortality rate.
The best growth result was obtained with a 10 m ditch spacing and strong fertilization. As it is difficult to decrease the 10 m ditch spacing for cost reasons, it can be concluded that on such oligotrophic peatlands as were used in this experiment, only an average growth level in the seedling stands can be reached even with the most efficient forest improvement measures. Broadcast fertilization used in the experiment, at least in large doses, increases seedling mortality, as well as the coverage of the ground vegetation, particularly that of cottongrass and fireweed, and also the shrub height, thus increasing competition. It cannot be recommended for afforestation, and today spot fertilization is used. According to this experiment natural seedlings seem once they have recovered after the first years, to grow better than the planted seedlings. This was true especially in the north and in areas, where drainage was not efficient. The height and height growth of the seedlings were to a large extent dependent on the temperature sum.
The PDF includes a summary in English.