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Silva Fennica 1926-1997
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Acta Forestalia Fennica
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1913-1932

Articles by Timo Pukkala

Category : Article

article id 5636, category Article
Timo Pukkala, Jyrki Kangas, Matleena Kniivilä, Anne-Mari Tiainen. (1997). Integrating forest-level and compartment-level indices of species diversity with numerical forest planning. Silva Fennica vol. 31 no. 4 article id 5636. https://doi.org/10.14214/sf.a8538
Keywords: simulation; heuristics; biodiversity conservation; forestry decision-making; environmental planning
Abstract | View details | Full text in PDF | Author Info

The study proposes a technique which enables the computation of user-defined indices for species diversity. These indices are derived from characteristics, called diversity indicators, of inventory plots, stand compartments, and the whole forest holding. The study discusses the modifications required to be made to typical forest planning systems due to this kind of biodiversity computation. A case study illustrating the use of the indices and a modified forest planning system is provided. In the case study, forest-level species diversity index was computed from the volume of dead wood, volume of broadleaved trees, area of old forest, and between-stand variety.

At the stand level, the area of old forest was replaced by stand age, and variety was described by within-stand variety. All but one of the indicators were further partitioned into two to four sub-indicators. For example, the volume of broadleaved trees was divided into volumes of birch, aspen, willow, and other tree species. The partial contribution of an indicator to the diversity index was obtained from a sub-priority function, determined separately for each indicator. The diversity index was obtained when the partial contributions were multiplied by the weights of the corresponding indicators and then were summed. The production frontiers computed for the harvested volume and diversity indices were concave, especially for the forest-level diversity index, indicating that diversity can be maintained at satisfactory level with medium harvest levels.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Kangas, E-mail: jk@mm.unknown
  • Kniivilä, E-mail: mk@mm.unknown
  • Tiainen, E-mail: at@mm.unknown
article id 5519, category Article
Jyrki Kangas, Jari Karsikko, Laura Laasonen, Timo Pukkala. (1993). A method for estimating the suitability function of wildlife habitat for forest planning on the basis of expertise. Silva Fennica vol. 27 no. 4 article id 5519. https://doi.org/10.14214/sf.a15680
Keywords: forest management; models; wildlife management; black grouse; production functions; multiple use; Lururus tetrix
Abstract | View details | Full text in PDF | Author Info

In the method presented in this study, a group of experts evaluate, in a pairwise manner, a set of forest areas with respect to the game species considered. On the basis of these comparisons, relative priorities of forest areas are estimated using the eigenvalue technique. Using regression analysis, a habitat suitability function is estimated in which the priority is predicted by measures already familiar in forest planning. As a case study, a habitat suitability function was estimated for black grouse (Tetrao tetrix, Lururus tetrix L.). The function is applicable in forestry planning carried out using modern planning techniques.

The PDF includes an abstract in Finnish.

  • Kangas, E-mail: jk@mm.unknown (email)
  • Karsikko, E-mail: jk@mm.unknown
  • Laasonen, E-mail: ll@mm.unknown
  • Pukkala, E-mail: tp@mm.unknown
article id 5511, category Article
Veli Pohjonen, Timo Pukkala. (1993). Cupressus lusitanican tuotos Etiopiassa. Silva Fennica vol. 27 no. 3 article id 5511. https://doi.org/10.14214/sf.a15672
English title: Yield of Cupressus lusitanica in Ethiopia.
Keywords: distribution; plantation; diameter; yield, models; yield tables; Cupressus lusitanica; thinning regimes
Abstract | View details | Full text in PDF | Author Info

Yield of Cupressus lusitanica Mill. was modelled by predicting the diameter distribution of trees at given stand ages. The beta distribution was used as a theoretical distribution. The models used for the calculation of diameter distribution were based on 66 temporal sample plots with varying age, site and stand density. The growing sites of Cupressus lusitanica were divided into four classes on the basis of age and dominant height. Using the stand models developed in the study, the yield and profitability of different thinning schedules was evaluated by a simulation technique. In the simulated treatment regimes, the mean annual increment varied from 6.6 m3/ha in the poorest site class to 16.6 m3/ha in the best class with rotation lengths ranging from 25 years (best sites) to 34 years (poorest sites). With typical planting densities (1,600 trees/ha), thinnings increase the total harvest by a few percentage points and improved the profitability of plantation forestry.
The PDF includes an abstract in Finnish.

  • Pohjonen, E-mail: vp@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5492, category Article
Ismo Nousiainen, Timo Pukkala. (1992). Use of computer graphics for predicting the amenity of forest trails. Silva Fennica vol. 26 no. 4 article id 5492. https://doi.org/10.14214/sf.a15653
Keywords: forest management; forest planning; simulation; landscape; recreation; amenity value of forests
Abstract | View details | Full text in PDF | Author Info

Ten trails, one kilometre each, were evaluated by 15 persons for scenic beauty, recreational value and variety. All trails passed through commercially managed forests dominated by conifers. The trails were first evaluated by viewing computer simulations based on a series of graphical illustrations of forest landscapes, then from a slide show, and finally in the field. In the computer simulation and slide show, landscape pictures along the trail at an interval of 35–40 m were presented for 3–4 seconds. The ranks between slide show and field were slightly more similar than those between simulation and field. The mean correlation of 12 persons between the field ranking and assessment of either computer simulations or slide shows or graphics than scenic beauty or recreational value. Spearman’s rank correlations computed from median scores of a group of 12 peers were clearly better than the average of individual persons varying from 0.6 to 0.9.

The PDF includes an abstract in Finnish

  • Nousiainen, E-mail: in@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5490, category Article
Timo Pukkala, Jari Karsikko, Taneli Kolström. (1992). A spatial model for the diameter of thickest branch of Scots pine. Silva Fennica vol. 26 no. 4 article id 5490. https://doi.org/10.14214/sf.a15651
Keywords: Pinus sylvestris; simulation; timber quality; competition; stand density; spatial pattern
Abstract | View details | Full text in PDF | Author Info

The model predicts the base diameter of the thickest living branch of a tree growing in a planted or naturally regenerated even-aged stand. A mixed model type was used in which the residual variation was divided into within-stand and between-stand components. The study material consisted of 779 trees measured in 12 plots located in 20 to 35 years old Scots pine (Pinus sylvestris L.) stands (breast height age 10 to 20 years). Branch diameter was closely connected to the breast height diameter of the stem. In a stand of a certain age, competition by close neighbours slightly decreased branch diameter in a given diameter class. According to the model, the greatest difference is between trees subjected to very little competition and those subjected to normal competition. The model was used in simulated stands with varying age, density, and tree arrangement. The simulations showed that trees with rapid diameter growth at young age had thicker branches at a given breast height diameter than trees with slower diameter growth. However, a very slow growth rate did not produce trees with branches thinner than those possessing a medium growth rate.
The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Karsikko, E-mail: jk@mm.unknown
  • Kolström, E-mail: tk@mm.unknown
article id 5484, category Article
Jyrki Kangas, Timo Pukkala. (1992). A decision theoretic approach applied to goal programming of forest management. Silva Fennica vol. 26 no. 3 article id 5484. https://doi.org/10.14214/sf.a15645
Keywords: forest management; models; forest planning; decision analysis; goal programming; optimization methods
Abstract | View details | Full text in PDF | Author Info

An alternative approach to formulating a forestry goal programming problem is presented. First, single objective optima levels are solved. The Analytical Hierarchy Process is applied in the estimation of a priori weights of deviations from the goal target levels. The ratios of the weights can be interpreted as relative importance of the goals, respectively. The sum of the weighted deviations from all single optima levels associated with the management goals is minimized. Instead of absolute deviations, relative ones are used. A case study problem of forest management planning with several objectives, measured in different units, is analysed.

The PDF includes an abstract in Finnish.

  • Kangas, E-mail: jk@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5449, category Article
Timo Pukkala, Taneli Kolström. (1991). Effect of spatial pattern of trees on the growth of Norway spruce stand. Silva Fennica vol. 25 no. 3 article id 5449. https://doi.org/10.14214/sf.a15603
Keywords: Picea abies; thinnings; growth; spatial distribution; simulation models; biological competition
Abstract | View details | Full text in PDF | Author Info

The simulation model consists of a method to generate theoretical Norway spruce (Picea abies (L.) H. Karst.) stands, and a spatial growth model to predict the growth of these stands. The stand generation procedure first predicts the tree diameters from a few stand characteristics and from tree locations. Tree age and height are predicted using spatial models. Spatial growth models were made for both diameter growth and basal area growth. Past growth was used as a predictor in one pair of models and omitted in another pair. The stand generation method and the growth models were utilized in studying the effect of tree arrangement and thinning method on the growth of a Norway spruce stand.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Kolström, E-mail: tk@mm.unknown
article id 5424, category Article
Timo Pukkala, Veli Pohjonen. (1990). Use of linear programming in land use planning in the Ethiopian highlands. Silva Fennica vol. 24 no. 2 article id 5424. https://doi.org/10.14214/sf.a15578
Keywords: plantations; fuelwood; mathematical models; grazing; Eucalyptus globulus; arable crops
Abstract | View details | Full text in PDF | Author Info

Linear programming was used to analyse the land use alternatives in the Debre Birhan Fuelwood Plantation area, in the central highlands of Ethiopia. The region represents a rural, high-altitude area, where the main land uses are grazing and cultivation of barley, wheat and pulses. To alleviate fuelwood shortage, large plantations of Eucalyptus globulus Labill. have been established. Livestock has traditionally used the major part of the production capacity of the sites. A decrease in the number of cattle would facilitate a considerable increase in the production of cereals, pulses, fuelwood and construction timber. The optimal share of the land for arable crops, grazing and tree plantations would be about 40, 45 and 15% respectively.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Pohjonen, E-mail: vp@mm.unknown
article id 5419, category Article
Timo Pukkala, Owen Mubita, Jussi Saramäki. (1990). Management planning system for tree plantations. Silva Fennica vol. 24 no. 2 article id 5419. https://doi.org/10.14214/sf.a15573
Keywords: diameter distribution; plantations; yield model; Zambia; simulation model; Pinus kesiya; timber management planning; multiobjective optimization
Abstract | View details | Full text in PDF | Author Info

The paper presents a simple model of long-term forest management planning in tree plantations. The model is particularly suitable for developing countries where the research resources are limited. The management plan is prepared in two steps. First, one or several treatment schedules are simulated for each calculation unit (age class, compartment, etc.) over the selected planning period. Second, an optimal combination treatment schedules according to the selected objectives and constraints is searched by mathematical programming. The simulation of growth is based on the prediction of the diameter distribution at the desired time point. All stand characteristics are derived from this distribution. The models needed in the yield simulation can be estimated from temporary sample plots. A case study management plan for 13,000 ha of Pinus kesiya (Royle ex Gordon) plantations in Zambia is presented to demonstrate the system.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Mubita, E-mail: om@mm.unknown
  • Saramäki, E-mail: js@mm.unknown
article id 5379, category Article
Timo Kuuluvainen, Timo Pukkala. (1989). Effect of Scots pine seed trees on the density of ground vegetation and tree seedlings. Silva Fennica vol. 23 no. 2 article id 5379. https://doi.org/10.14214/sf.a15536
Keywords: Pinus sylvestris; natural regeneration; spatial variation; ground vegetation; seedlings; seed trees; regeneration models; ecological fields; resource consumption; competitive interface
Abstract | View details | Full text in PDF | Author Info

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5376, category Article
Timo Pukkala. (1989). Predicting diameter growth in even-aged Scots pine stands with a spatial and non-spatial model. Silva Fennica vol. 23 no. 2 article id 5376. https://doi.org/10.14214/sf.a15533
Keywords: Pinus sylvestris; growth prediction; spatial distribution; growth models; tree models
Abstract | View details | Full text in PDF | Author Info

The single tree growth models presented in this study were based on about 4,000 trees measured in 50 even-aged Scots pine (Pinus sylvestris L.) sample plots with varying density, spatial pattern of trees and stand age. Predictors that used information about tree locations decreased the relative standard error of estimate by 10 percentage points (15%), if past growth was not used as a predictor, and about 15 percentage points (30%) when past growth was one of the predictors. When ranked according to the degree of determination, the best growth models were obtained for the basal area increment, the next best for relative growth, and the poorest for diameter increment. The past growth decreased the relative standard error of estimate by 15–20 percentage points, but did not make the spatial predictors unnecessary. The degree of determination of the spatial basal area growth model was almost 80% if the past growth was unknown and almost 90% if the past growth was known. Variables that described the amount of removed competition did not improve the growth models.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5375, category Article
Timo Pukkala. (1989). Prediction of tree diameter and height in a Scots pine stand as a function of the spatial pattern of trees. Silva Fennica vol. 23 no. 2 article id 5375. https://doi.org/10.14214/sf.a15532
Keywords: Pinus sylvestris; diameter distribution; competition; spatial distribution; simulations studies; tree models
Abstract | View details | Full text in PDF | Author Info

The study presents two methods of predicting tree dimensions in a Scots pine (Pinus sylvestris L.) stand if only the location of trees is known. The first method predicts the tree diameter from the spatial location of neighbours. In the second method the diameter distribution of a subarea is estimated from the local stand density. This distribution is then sampled to obtain diameters. In both methods the tree height is predicted with a spatial model on the basis of diameters and locations of trees. The main purpose of the presented models is to generate realistic stands for simulation studies.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5363, category Article
Veli Pohjonen, Timo Pukkala. (1988). Profitability of establishing Eucalyptus globulus plantations in the Central Highlands of Ethiopia. Silva Fennica vol. 22 no. 4 article id 5363. https://doi.org/10.14214/sf.a15520
Keywords: reforestation; Ethiopia; Eucalyptus globulus; simulation of growth; economic analysis; land use planning
Abstract | View details | Full text in PDF | Author Info

The economic analysis is based on computer simulations which covered a seedling rotation and three successive coppice rotations. Calculations were carried out for the four site productivity classes in Eucalyptus globulus plantations. The rotation length that maximized the land expectation value is 12–20 years for seedling rotation and 8–16 years for coppice rotations with discounting rates 2–8%. The mean wood production is over 40 m3/ha/a in the best site class and about 10 m3/ha/a in the poorest class with rotation lengths ranging from 10 to over 20 years. Thinnings increase the wood production and land expectation value by a few percentage points. In areas suitable to Eucalyptus globulus growth, the land expectation value is considerably higher in forestry than in agriculture, except in very poor areas or with very high rate of interest.

The PDF includes an abstract in Finnish.

  • Pohjonen, E-mail: vp@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5346, category Article
Timo Pukkala. (1988). Methods to incorporate the amenity of landscape into forest management planning. Silva Fennica vol. 22 no. 2 article id 5346. https://doi.org/10.14214/sf.a15503
Keywords: forest management; landscape management; multipurpose forestry; multiobjective planning; computer simulation; mathematical optimization; management plan
Abstract | View details | Full text in PDF | Author Info

The study presents methods that incorporate the amenity of a forest area into the management planning. The management plan is based on treatment schedules simulated for each compartment over the 20-year planning period. The best combination of treatment schedules is selected by multi-objective optimization. The amenity is divided into two parts: (1) within-stand amenity and (2) the amenity of landscape when viewed afar (distant scene). The within-stand amenity is expressed in terms of adjective sum which is estimated from stand characteristics. The adjective sum of the whole area in a selected year can be taken as an objective or constraining variable of optimization. The assessment of the distant scene is based on computer illustrations which show the predicted temporal change of landscape according to a particular management plan.

The PDF includes a summary in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5338, category Article
Timo Pukkala. (1988). Effect of spatial distribution of trees on the volume increment of a young Scots pine stand. Silva Fennica vol. 22 no. 1 article id 5338. https://doi.org/10.14214/sf.a15495
Keywords: Pinus sylvestris; simulation; competition; spatial distribution; growth model; spatial pattern
Abstract | View details | Full text in PDF | Author Info

The effect of grouping on 5-year old volume increment was studied by a simulation technique using spatial growth models estimated in Scots pine (Pinus sylvestris L.) stands in the phase of the first commercial thinning. A total of 24 model stands were regenerated by applying 12 spatial processes for two different diameter distributions. In addition to model stands, 6 different thinnings were simulated in two real stands. The clustering of trees was described with Fisher’s grouping index and by estimating the relative interception of diffuse radiation. In model stands with constant diameter distribution the correlation between the grouping index and volume increment ranged from -0.81 to -0.91. The correlation between volume increment and interception was 0.81–0.83 with one diameter distribution and 0.70 if both distributions were combined. In one thinned stand the correlation between the growth estimate and grouping index varied between -0.33 and 0.76. The correlation between interception and growth was about 0.30 in one stand and 0.72 if both stands were combined. Small irregularities do not decrease the volume production of a young Scots pine stand, but if the clustering is considerable or there are reasonably wide harvest strips, growth will be reduced by 10–20%.

The PDF includes a summary in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5316, category Article
Timo Pukkala, Timo Kuuluvainen. (1987). Effect of canopy structure on the diurnal interception of direct solar radiation and photosynthesis in a tree stand. Silva Fennica vol. 21 no. 3 article id 5316. https://doi.org/10.14214/sf.a15473
Keywords: canopy; simulation; shading; crown type; interception; tree ideotype; radiation regime
Abstract | View details | Full text in PDF | Author Info

The utilization of direct radiation was studied in five model stands of Poisson-type tree distribution and cone-shaped crowns. The radiation extinction depended on the self-shading of the crown and the shading caused by other trees. The results indicate that at low sun elevation a stand populated by very narrow-crowned trees is most effective in light interception and photosynthesis. At high sun elevation a broad-crowned canopy is best illuminated and most favourable for photosynthesis. A stand with a two-storey canopy is effective in all latitudes when the crowns are moderately narrow. In two-storey canopies the foliage of the lower storey can be better illuminated than in the lower parts of the upper storey, because of the smaller self-shading in the small crowns of the lower storey. A canopy where the crown volume is concentrated on few big crowns is less effective than a canopy consisting of many small crowns.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Kuuluvainen, E-mail: tk@mm.unknown
article id 5312, category Article
Timo Pukkala. (1987). Siementuotannon vaikutus kuusen ja männyn vuotuiseen kasvuun. Silva Fennica vol. 21 no. 2 article id 5312. https://doi.org/10.14214/sf.a15469
English title: Effect of seed production on the annual growth of Picea abies and Pinus sylvestris.
Original keywords: kuusi; mänty; Lappi; siemensato; sädekasvu; kesäpuu
English keywords: Pinus sylvestris; Norway spruce; Picea abies; Scots pine; Finland; latewood; Lapland; radial growth; seed crop
Abstract | View details | Full text in PDF | Author Info

The study material consisted of 13 rather old Norway spruce (Picea abies (L.) H. Karst.) and 17 Scots pine (Pinus sylvestris L.) stands located in different parts of Finland. In each stand the seed crops, radial growth and amount of latewood were measured during a period of about ten years. Seed production reduces the radial growth of spruce and pine in the year of seed maturing. In Southern and Central Finland also the proportion of latewood is reduced. Seed production accounts for about 14% of the variation in radial growth of a spruce stand growing in Lapland, and 27% in other parts of Finland. In pine stands the seed crop explains 19% of the variation in radial growth in Lapland, and only 7% in the rest of Finland. In spruce stands an average seed crop reduces radial growth by 14% in Lapland and 5% in the rest of the country. An abundant seed production causes a reduction of about 20%. In southern parts of Finland, the proportion of latewood is reduced by 5% in an average seed year and by 24% in a good seed year. In pine stands an average seed crop decreases the width of annual ring by 5%, and a good seed crop by 15%. Outside Lapland, also the proportion of latewood is reduced: in an average seed year by 5%, and in a good seed year by 16%. The reduction in volume growth of spruce stands due to an average seed crop was estimated to be about 10% in Lapland, and 6% in other parts of Finland. A prolific seed production causes a reduction of 20%. In old pine stands the reduction is 5% in an average seed year, and 15% in a good seed year.

The PDF includes an abstract in English.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5311, category Article
Timo Pukkala. (1987). Kuusen ja männyn siemensadon ennustemalli. Silva Fennica vol. 21 no. 2 article id 5311. https://doi.org/10.14214/sf.a15468
English title: Model for predicting the seed crop of Picea abies and Pinus sylvestris.
Original keywords: kuusi; mänty; luontainen uudistaminen; Lappi; siemensato; lämpötila; kukinnan aloitus; ennustemalli
English keywords: Pinus sylvestris; Norway spruce; Picea abies; natural regeneration; Scots pine; Finland; Lapland; predictive modelling; seed crop; flowering initiation; influence of temperature
Abstract | View details | Full text in PDF | Author Info

The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.

In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.

In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.

The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.

The PDF includes an abstract in English.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5306, category Article
Timo Pukkala, Taneli Kolström. (1987). Competition indices and the prediction of radial growth in Scots pine. Silva Fennica vol. 21 no. 1 article id 5306. https://doi.org/10.14214/sf.a15463
Keywords: Pinus sylvestris; stand characteristics; simulation; tree growth; competition; radial growth
Abstract | View details | Full text in PDF | Author Info

The effect of competition on the radial growth of Scots pine (Pinus sylvestris L.) was studied in three naturally regenerated stands located in North Karelia, Finland. The competition situation of an individual tree was described with various competition indices which depended on the sizes and distances from the neighbouring trees. One competition index explained about 50% of the variation in 5-year radial growth in one stand. If all stands were combined, one index explained 43.5%, two indices 48.9% and three indices 51.2% of the variation. In one stand, the best competition indices accounted for about 20% of that variation which could not be explained by tree diameter. If all three stands were combined, the best index explained 11% of the residual variation. About 40% of the variation in 5-year radial growth could not be explained by the diameter and competition indices.
The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
  • Kolström, E-mail: tk@mm.unknown
article id 5305, category Article
Timo Pukkala. (1987). Simulation model for natural regeneration of Pinus sylvestris, Picea abies, Betula pendula and Betula pubescens. Silva Fennica vol. 21 no. 1 article id 5305. https://doi.org/10.14214/sf.a15462
Keywords: Pinus sylvestris; Betula pendula; Picea abies; germination; Betula pubescens; mortality; seed crop; computer model; stochastic simulation; birth model
Abstract | View details | Full text in PDF | Author Info

In the model the regeneration process is derived into three subprocesses: birth, growth and mortality of seedlings. The main emphasis is on the birth process where the following phases are simulated: seed crop, quality of seeds, maturity of seeds, predation of seeds and germination. The parameters are based on data published in Finland. Part of the parameters are obtained directly from the investigations and part is proposed by the author. The model can be calibrated by changing parameter values. The simulation is made with the help of random numbers which have the same means as the estimates and the same distributions as the residuals of the equations used in simulation. The time step of the model is one year. The number of emerged seedlings in one year is obtained by multiplying the seed crop with the probabilities that the seed passes different phases of the birth process. Because of stochasticity the regeneration period is simulated several times. From the results it is possible to evaluate the risk and succeeding probability of the regeneration. The main drawbacks of the simulation method are the lack of empirical parameters and the difficulty of testing. The model could be further developed by including spatiality into the model.

The PDF includes an abstract in Finnish.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5302, category Article
Seppo Kellomäki, Heikki Hänninen, Taneli Kolström, Ahti Kotisaari, Timo Pukkala. (1987). A tentative model for describing the effects of some regenerative process on the properties of natural seedling stands. Silva Fennica vol. 21 no. 1 article id 5302. https://doi.org/10.14214/sf.a15459
Keywords: Pinus sylvestris; natural regeneration; Scots pine; modelling; stand density; seed crop; seed dispersal; parent tree
Abstract | View details | Full text in PDF | Author Info

The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.

As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.

The PDF includes an abstract in Finnish.

  • Kellomäki, E-mail: sk@mm.unknown (email)
  • Hänninen, E-mail: hh@mm.unknown
  • Kolström, E-mail: tk@mm.unknown
  • Kotisaari, E-mail: ak@mm.unknown
  • Pukkala, E-mail: tp@mm.unknown

Category : Article

article id 7513, category Article
Jyrki Kangas, Teppo Loikkanen, Timo Pukkala, Jouni Pykäläinen. (1996). A participatory approach to tactical forest planning. Acta Forestalia Fennica no. 251 article id 7513. https://doi.org/10.14214/aff.7513
Keywords: forest planning; public participation; optimization; heuristics; conflict management; decision analysis; participative planning
Abstract | View details | Full text in PDF | Author Info

The paper examines the needs, premises and criteria for effective public participation in tactical forest planning. A method for participatory forest planning utilizing the techniques of preference analysis, professional expertise and heuristic optimization is introduced. The techniques do not cover the whole process of participatory planning, but are applied as a tool constituting the numerical core for decision support. The complexity of multi-resource management is addressed by hierarchical decision analysis which assesses the public values, preferences and decision criteria toward the planning situation. An optimal management plan is sought using heuristic optimization. The plan can further be improved through mutual negotiations, if necessary. The use of the approach is demonstrated with an illustrative example. Its merits and challenges for participatory forest planning and decision making are discussed and a model for applying it in general forest planning context is depicted. By using the approach, valuable information can be obtained about public preferences and the effects of taking them into consideration on the choice of the combination of standwise treatment proposals for a forest area. Participatory forest planning calculations, carried out by the approach presented in the paper, can be utilized in conflict management and in developing compromises between competing interests.

  • Kangas, E-mail: jk@mm.unknown (email)
  • Loikkanen, E-mail: tl@mm.unknown
  • Pukkala, E-mail: tp@mm.unknown
  • Pykäläinen, E-mail: jp@mm.unknown

Category : Research article

article id 10732, category Research article
Ana Aza, A. Maarit I. Kallio, Timo Pukkala, Ari Hietala, Terje Gobakken, Rasmus Astrup. (2022). Species selection in areas subjected to risk of root and butt rot: applying Precision forestry in Norway. Silva Fennica vol. 56 no. 3 article id 10732. https://doi.org/10.14214/sf.10732
Keywords: Norway spruce; Scots pine; growth modelling; precision forestry; root and butt rot severity; tree species selection
Highlights: We present the best species to plant on previously spruce-dominated sites with different site indexes and rot levels; We recommend planting Norway spruce on low-rot sites, Scots pine on higher-rot sites, and allowing natural regeneration on low site indexes; We demonstrate the Precision forestry method for determining the optimal tree species in heterogenous stands; In the case study, the method increased net present value by approximately 6% on average.
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Norway’s most common tree species, Picea abies (L.) Karst. (Norway spruce), is often infected with Heterobasidion parviporum Niemelä & Korhonen and Heterobasidion annosum (Fr.) Bref.. Because Pinus sylvestris L. (Scots pine) is less susceptible to rot, it is worth considering if converting rot-infested spruce stands to pine improves economic performance. We examined the economically optimal choice between planting Norway spruce and Scots pine for previously spruce-dominated clear-cut sites of different site indexes with initial rot levels varying from 0% to 100% of stumps on the site. While it is optimal to continue to plant Norway spruce in regions with low rot levels, shifting to Scots pine pays off when rot levels get higher. The threshold rot level for changing from Norway spruce to Scots pine increases with the site index. We present a case study demonstrating a practical method (“Precision forestry”) for determining the tree species in a stand at the pixel level when the stand is heterogeneous both in site indexes and rot levels. This method is consistent with the concept of Precision forestry, which aims to plan and execute site-specific forest management activities to improve the quality of wood products while minimising waste, increasing profits, and maintaining environmental quality. The material for the study includes data on rot levels and site indexes in 71 clear-cut stands. Compared to planting the entire stand with a single species, pixel-level optimised species selection increases the net present value in almost every stand, with average increase of approximately 6%.

  • Aza, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway ORCID https://orcid.org/0000-0002-6416-6697 E-mail: anfe@nmbu.no (email)
  • Kallio, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: maarit.kallio@nmbu.no
  • Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
  • Hietala, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: ari.hietala@nibio.no
  • Gobakken, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: terje.gobakken@nmbu.no
  • Astrup, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: rasmus.astrup@nibio.no
article id 10370, category Research article
Juha Lappi, Timo Pukkala. (2020). Analyzing ingrowth using zero-inflated negative binomial models. Silva Fennica vol. 54 no. 4 article id 10370. https://doi.org/10.14214/sf.10370
Keywords: regeneration; continuous cover forestry; count data; generalized linear model; overdispersion; right-censoring
Highlights: Models were developed to describe ingrowth in national forest inventory data; The data were more dispersed than Poisson data and included many zeros; Fixed-effects models had larger zero-inflation probability and overdispersion parameter than mixed-effect models; Mixed-effects models had larger likelihood than fixed-effects models but provided biased predictions; Prediction of right-censored ingrowth may be useful owing to large overdispersion.
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Ingrowth is an important element of stand dynamics in several silvicultural systems, especially in continuous cover forestry. Earlier predictive models for ingrowth in Finnish forests are few and not based on up-to-date statistical methods. Ingrowth is here defined as the number of trees over 1.3 m entering a plot. This study developed new ingrowth models for Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.) and birch (Betula pendula Roth and B. pubescens Ehrh.) using data from the permanent sample plots of the Finnish national forest inventory. The data were over-dispersed compared to a Poisson process and had many zeros. Therefore, a zero-inflated negative binomial model was used. The total and species-specific stand basal areas, temperature sum and fertility class were used as predictors in the ingrowth models. Both fixed-effects and mixed-effects models were fitted. The mixed-effects model versions included random plot effects. The mixed-effects models had larger likelihoods but provided biased predictions. Also censored prediction was considered where only a certain maximum number of ingrowth trees were accepted for a plot. The models predicted most pine ingrowth in pine-dominated stands on sub-xeric and xeric sites where stand basal area was low. The predicted amount of spruce ingrowth was maximized when the basal area of spruce was 13 m2 ha–1. Increasing temperature sum increased spruce ingrowth. Predicted birch ingrowth decreased with increasing stand basal area and towards low fertility classes. An admixture of pine increased the predicted amount of spruce ingrowth.

  • Lappi, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: juha.lappi.sjk@gmail.com (email)
  • Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
article id 10217, category Research article
Xingji Jin, Timo Pukkala, Fengri Li, Lihu Dong. (2019). Developing growth models for tree plantations using inadequate data – a case for Korean pine in Northeast China. Silva Fennica vol. 53 no. 4 article id 10217. https://doi.org/10.14214/sf.10217
Keywords: Pinus koraiensis; optimization-based modeling; quantile regression; self-thinning
Highlights: The permanent sample plots of Chinese plantation trees have not been designed for producing data for growth modeling; We used various methods to deal with the inadequacies of sample plot data; Optimization was used to fit diameter increment and survival models using data with varying measurement intervals and tree identification errors; Quantile regression was used to model self-thinning limit.
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Korean pine (Pinus koraiensis Siebold & Zucc.) is economically the most important tree species in northeast China. Korean pine plantations are established and managed for the production of timber and seeds. Despite the importance of the species, few models have been developed for the comparison of alternative management schedules. Model development is affected by the fact that permanent sample plots and thinning experiments have not been designed and managed for modeling purposes. The permanent sample plots include few non-thinned plots, and weak trees are removed in thinning treatments, leading to low mortality rate. Moreover, the measurement interval is irregular. This study used optimization-based modeling approach in tree-level diameter increment and survival modeling to deal with the above problems. Models for self-thinning limit were developed to alleviate the problem of underestimated mortality arising from the features of the data. In addition, improved site index and individual-tree height models were developed. The model of Lundqvist and Korf was used as the site index model and the model proposed by Schumacher as the height model. Quantile regression was used to model the maximum stand basal area and maximum number of trees as a function of mean tree diameter and site index. Tree diameter, stand basal area, basal area in larger trees and site index were used as the predictors of diameter increment and tree survival. The models developed in this study constitute a model set that is suitable for simulation and optimization studies. The models produced simulation results that correspond to measured stand development.

  • Jin, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0003-2971-2709 E-mail: xingji_jin@163.com
  • Pukkala, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China; University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi (email)
  • Li, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0002-4058-769X E-mail: fengrili@126.com
  • Dong, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0002-3985-9475 E-mail: ldonglihu2006@163.com
article id 10187, category Research article
Timo Pukkala, Kjersti Holt Hanssen, Kjell Andreassen. (2019). Stem taper and bark functions for Norway spruce in Norway. Silva Fennica vol. 53 no. 3 article id 10187. https://doi.org/10.14214/sf.10187
Keywords: forest management; Picea abies; Kozak model; variable-exponent taper function
Highlights: New variable-exponent stem taper and bark functions were developed for Norway spruce; Both fixed and mixed-effects models were developed; Site index and tree age had statistically significant but small effects on stem taper.
Abstract | Full text in HTML | Full text in PDF | Author Info

Based on data from long-term experimental fields with Norway spruce (Picea abies (L.) H. Karst.), we developed new stem taper and bark functions for Norway. Data was collected from 477 trees in stands across Norway. Three candidate functions which have shown good performance in previous studies (Kozak 02, Kozak 97 and Bi) were fitted to the data as fixed-effects models. The function with the smallest Akaike Information Criterion (AIC) was then chosen for additional analyses, fitting 1) site index-dependent and 2) age-dependent versions of the model, and 3) fitting a mixed-effects model with tree-specific random parameters. Kozak 97 was found to be the function with the smallest AIC, but all three tested taper functions resulted in fairly similar predictions of stem taper. The site index-dependent function reduced AIC and residual standard error and showed that the effect of site index on stem taper is different in small and large trees. The predictions of the age-independent and age-dependent models were very close to each other. Adding tree-specific random parameters to the model clearly reduced AIC and residual variation. However, the results suggest that the mixed-effects model should be used only when it is possible to calibrate it for each tree, otherwise the fixed-effects Kozak 97 model should be used. A model for double bark thickness was also fitted as fixed-effects Kozak 97 model. The model behaved logically, predicting larger relative but smaller absolute bark thickness for small trees.

article id 131, category Research article
Timo Pukkala, Tatu Hokkanen, Teijo Nikkanen. (2010). Prediction models for the annual seed crop of Norway spruce and Scots pine in Finland. Silva Fennica vol. 44 no. 4 article id 131. https://doi.org/10.14214/sf.131
Keywords: flowering; cone crops; seed years; mixed model
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Many studies indicate that the flowering abundance of boreal trees strongly correlates with the weather conditions of the previous summer. This study developed prediction models for the seed crops of Norway spruce and Scots pine using weather variables one and two years prior to flowering year as predictors. Weather data, systematically recorded at many weather stations, were obtained from the Finnish Meteorological Institute. Seed crop monitoring data came from 22 spruce stands and 44 pine stands. In every stand, seed crop has been monitored for many years, the longest continuous period being 45 years. Monthly mean temperatures, monthly rainfalls, and periodical temperature sums were used as predictors in the seed crop models. Generally, both tree species flowered abundantly one year after a warm summer and two years after a cool summer. While the models only explained about 45% of the variation in the annual seed crop, they accurately predicted good and bad seed years: when the models predicted good seed crops the likelihood to have at least a medium seed crop was very high and when the models predicted small seed crops, the likelihood to obtain medium or good seed crop was very low. Therefore, the models reliably predict if a particular year will be a good seed year or a poor seed year. These predictions can be used in forestry practice for proper timing of natural regeneration activities, and when activities in seed orchards are planned.
  • Pukkala, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi (email)
  • Hokkanen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: th@nn.fi
  • Nikkanen, Finnish Forest Research Institute, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: tn@nn.fi
article id 176, category Research article
María Pasalodos-Tato, Timo Pukkala, Antonio Rigueiro-Rodríguez, Esther Fernández-Nunez, María Rosa Mosquera-Losada. (2009). Optimal management of Pinus radiata silvopastoral systems established on abandoned agricultural land in Galicia (north-western Spain). Silva Fennica vol. 43 no. 5 article id 176. https://doi.org/10.14214/sf.176
Keywords: simulation; economic profitability; optimization; risk reduction; salvage
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Timber production has been the main objective in forest production in Galicia for a long time. Nevertheless, factors such as fire risk and the need to obtain non-timber benefits make other production alternatives like silvopastoral systems worth of consideration. Integration of grazing in the production system not only diversifies products and benefits, but also decreases fire risk by enhancing fuel control. Nonetheless, few studies have examined the economic profitability of these systems. This article analyses the economics of silvopastoral systems established on abandoned agricultural soils afforested with Pinus radiata D. Don. Different tree planting densities, discounting rates, grass values and fire risk scenarios were analysed. The technique employed is based on the combination of an optimization algorithm and a simulator of stand growth and grass yield. The most profitable schedules were obtained with initial stand densities of 1500 trees per hectare. However, with high unit values of pasture production (high value of grass), schedules with an initial stand density of 500 trees per hectare were the most profitable. When the risk of fire was included in the analyses, silvopastoral systems were always more profitable than timber production systems. With an assumption that grazing reduces fire risk thinnings should be done earlier and heavier to reduce the expected losses due to fire and to promote grass production. This lengthens the pasture period. In general, rotation lengt
  • Pasalodos-Tato, INIA, Instituto Nacional de Investigación y Tecnología Agraria y Alimentaria. Madrid, Spain E-mail: pasalodos.maria@inia.es (email)
  • Pukkala, University of East Finland, Joensuu, Finland E-mail: tp@nn.fi
  • Rigueiro-Rodríguez, University of Santiago de Compostela, Lugo, Spain E-mail: arr@nn.es
  • Fernández-Nunez, University of Santiago de Compostela, Lugo, Spain E-mail: efn@nn.es
  • Mosquera-Losada, University of Santiago de Compostela, Lugo, Spain E-mail: mrml@nn.es
article id 211, category Research article
Timo Pukkala. (2009). Population-based methods in the optimization of stand management. Silva Fennica vol. 43 no. 2 article id 211. https://doi.org/10.14214/sf.211
Keywords: Particle Swarm Optimization; differential evolution; evolution strategy; Nelder-Mead method; amoeba search; polytope search; evolutionary computing
Abstract | View details | Full text in PDF | Author Info
In Finland, the growth and yield models for tree stands are simulation programs that consist of several sub-models. These models are often non-smooth and non-differentiable. Direct search methods such as the Hooke-Jeeves algorithm (HJ) are suitable tools for optimizing stand management with this kind of complicated models. This study tested a new class of direct search methods, namely population-based methods, in the optimization of stand management. The tested methods were differential evolution, particle swarm optimization, evolution strategy, and the Nelder-Mead method. All these methods operate with a population of solution vectors, which are recombined and mutated to obtain new candidate solutions. The management schedule of 719 stands was optimized with all population-based methods and with the HJ method. The population-based methods were competitive with the HJ method, producing 0.57% to 1.74% higher mean objective function values than HJ. On the average, differential evolution was the best method, followed by particle swarm optimization, evolution strategy, and Nelder-Mead method. However, differences between the methods were small, and each method was the best in several stands. HJ was alone the best method in 7% of stands, and a population based method in 3% (Nelder-Mead) to 29% (differential evolution) of stands. All five methods found the same solution in 18% of stands.
  • Pukkala, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
article id 299, category Research article
Hongcheng Zeng, Timo Pukkala, Heli Peltola, Seppo Kellomäki. (2007). Application of ant colony optimization for the risk management of wind damage in forest planning. Silva Fennica vol. 41 no. 2 article id 299. https://doi.org/10.14214/sf.299
Keywords: harvest scheduling; heuristics; genetic algorithms; simulated annealing; spatial optimization
Abstract | View details | Full text in PDF | Author Info
Ant colony optimization (ACO) is still quite a new technique and seldom used in the field of forest planning compared to other heuristics such as simulated annealing and genetic algorithms. This work was aimed at evaluating the suitability of ACO for optimizing the clear-cut patterns of a forest landscape when aiming at simultaneously minimizing the risk of wind damage and maintaining sustainable and even flow of periodical harvests. For this purpose, the ACO was first revised and the algorithm was coded using the Visual Basic Application of the ArcGIS software. Thereafter, the performance of the modified ACO was demonstrated in a forest located in central Finland using a 30-year planning period. Its performance was compared to simulated annealing and a genetic algorithm. The revised ACO performed logically since the objective function value was improving and the algorithm was converging during the optimization process. The solutions maintained a quite even periodical harvesting timber while minimizing the risk of wind damage. Implementing the solution would result in smooth landscape in terms of stand height after the 30-year planning period. The algorithm is quite sensitive to the parameters controlling pheromone updating and schedule selecting. It is comparable in solution quality to simulated annealing and genetic algorithms.
  • Zeng, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: hongcheng.zeng@joensuu.fi (email)
  • Pukkala, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: tp@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
  • Kellomäki, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
article id 474, category Research article
Tero Heinonen, Mikko Kurttila, Timo Pukkala. (2007). Possibilities to aggregate raster cells through spatial optimization in forest planning. Silva Fennica vol. 41 no. 1 article id 474. https://doi.org/10.14214/sf.474
Keywords: threshold accepting; spatial optimisation; landscape metrics; old forest
Abstract | View details | Full text in PDF | Author Info
  • Heinonen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tero.heinonen@joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 332, category Research article
Marc Palahí, Timo Pukkala, Antoni Trasobares. (2006). Calibrating predicted tree diameter distributions in Catalonia, Spain. Silva Fennica vol. 40 no. 3 article id 332. https://doi.org/10.14214/sf.332
Keywords: calibration estimation; Weibull function; parameter prediction
Abstract | View details | Full text in PDF | Author Info
Several probability density functions have been used in describing the diameter distributions of forest stands. In a case where both the stand basal area and number of stems per hectare are assessed, the fitted or predicted distribution is scaled using only one of these variables, with the result that the distribution often gives incorrect values for the other variable. Using a distribution that provides incorrect values for known characteristics means wasting information. Calibrating the distribution so that it is compatible with the additional information on stand characteristics is a way to avoid such wasting. This study examined the effect of calibration on the accuracy of the predicted diameter distributions of the main tree species of Catalonia. The distributions were calibrated with and without considering the prediction errors of the frequencies of diameter classes. When prediction errors were assumed, the calibration was done with and without making allowance for estimation errors in the stand level calibration variables. Calibrated distributions were more accurate than non-calibrated in terms of sums of different powers of diameters. The set of calibration variables that gave the most accurate results included six stand variables: number of trees per hectare, stand basal area, basal-area-weighted mean diameter, non-weighted mean diameter, median diameter, and basal area median diameter. Of the tested three-variable combinations the best was: number of trees per hectare, stand basal area, and basal-area-weighted mean diameter. Means were more useful calibration variables than medians.
  • Palahí, Centre Tecnológic Forestal de Catalunya. Passeig Lluis Companys, 23, 08010, Barcelona, Spain E-mail: marc.palahi@ctfc.es (email)
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, 80101 Joensuu, Finland E-mail: tp@nn.fi
  • Trasobares, Foreco Technologies, Av. Diagonal 416, Estudio 2, Barcelona 08037, Spain E-mail: at@nn.es
article id 364, category Research article
Timo Pukkala, Jari Miina. (2005). Optimising the management of a heterogeneous stand. Silva Fennica vol. 39 no. 4 article id 364. https://doi.org/10.14214/sf.364
Keywords: Hooke and Jeeves algorithm; thinning schedules; within-stand variation
Abstract | View details | Full text in PDF | Author Info
The study presents a method for taking the heterogeneity of the stand into account in the optimisation of stand management. Heterogeneity refers to within-stand variation in stand density and/or other characteristics. A set of plots, corresponding to different sub-areas of the stand, represents the stand in calculations. Cuttings and other treatments of the plots are done simultaneously. The method was used to analyse how the optimal management depends on the heterogeneity of a Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) stand. The results supported the hypothesis that the heterogeneity of a stand decreases its optimal prior-thinning density. Also the remaining stand basal areas were lower in heterogeneous stands, especially in spruce. The effect of stand heterogeneity prior to the first commercial thinning still affected the timing of the second thinning, which had to be conducted earlier and at lower prior-thinning basal areas in heterogeneous stands. This happened despite the fact that the first thinning greatly decreased the within-stand variation in stand basal area. In addition, heterogeneity decreased the soil expectation value, net income and timber harvests.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
article id 396, category Research article
Timo Pukkala, Mikko Kurttila. (2005). Examining the performance of six heuristic optimisation techniques in different forest planning problems. Silva Fennica vol. 39 no. 1 article id 396. https://doi.org/10.14214/sf.396
Keywords: genetic algorithms; simulated annealing; ecological planning; habitat suitability index (HSI); Hero; random search; tabu search
Abstract | View details | Full text in PDF | Author Info
The existence of multiple decision-makers and goals, spatial and non-linear forest management objectives and the combinatorial nature of forest planning problems are reasons that support the use of heuristic optimisation algorithms in forest planning instead of the more traditional LP methods. A heuristic is a search algorithm that does not necessarily find the global optimum but it can produce relatively good solutions within reasonable time. The performance of different heuristics may vary depending on the complexity of the planning problem. This study tested six heuristic optimisation techniques in five different, increasingly difficult planning problems. The heuristics were evaluated with respect to the objective function value that the techniques were able to find, and the time they consumed in the optimisation process. The tested optimisation techniques were 1) random ascent (RA), 2) Hero sequential ascent technique (Hero), 3) simulated annealing (SA), 4) a hybrid of SA and Hero (SA+Hero), 5) tabu search (TS) and 6) genetic algorithm (GA). The results, calculated as averages of 100 repeated optimisations, were very similar for all heuristics with respect to the objective function value but the time consumption of the heuristics varied considerably. During the time the slowest techniques (SA or GA) required for convergence, the optimisation could have been repeated about 200 times with the fastest technique (Hero). The SA+Hero and SA techniques found the best solutions for non-spatial planning problems, while GA was the best in the most difficult problems. The results suggest that, especially in spatial planning problems, it is a benefit if the method performs more complicated moves than selecting one of the neighbouring solutions. It may also be beneficial to combine two or more heuristic techniques.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. BOX 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@forest.joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
article id 419, category Research article
Tero Heinonen, Timo Pukkala. (2004). A comparison of one- and two-compartment neighbourhoods in heuristic search with spatial forest management goals. Silva Fennica vol. 38 no. 3 article id 419. https://doi.org/10.14214/sf.419
Keywords: simulated annealing; Hero; tabu search; 2-optimal heuristic; spatial optimisation; random ascent
Abstract | View details | Full text in PDF | Author Info
This study presents a comparison of the performance of four heuristic techniques with one- and two-compartment neighbourhoods in harvest scheduling problems including a spatial objective variable. The tested heuristics were random ascent, Hero, simulated annealing and tabu search. All methods seek better solutions by inspecting the neighbourhood solutions, which are combinations that can be obtained by changing the treatment schedule in one (one-compartment neighbourhood) or two (two-compartment neighbourhood) compartments. The methods and neighbourhoods were examined in one artificial and four real landscapes ranging from 700 to 981 ha in size. The landscapes had 608 to 900 stand compartments, and the examined planning problems had 2986 to 4773 binary decision variables. The objective function was a multi-objective utility function. The spatial objective variable was the percentage of compartment boundary that joins two compartments, both of which are to be cut during the same 20-year period. The non-spatial objectives were net incomes of three consecutive 20-year management periods and the remaining growing stock volume at the end of the third 20-year period. In another problem formulation, the total harvest of the first 20-year period was used as an objective variable together with the spatial objective. The results showed that a two-compartment neighbourhood was systematically and often clearly better than a one-compartment neighbourhood. The improvements were greatest with the simplest heuristics, random ascent and Hero. Of the four heuristics, tabu search and simulated annealing proved to be the best methods, but with a two-compartment neighbourhood the differences between methods were negligible.
  • Heinonen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: th@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
article id 513, category Research article
Marjut Ihalainen, Kauko Salo, Timo Pukkala. (2003). Empirical prediction models for Vaccinium myrtillus and V. vitis-idaea berry yields in North Karelia, Finland. Silva Fennica vol. 37 no. 1 article id 513. https://doi.org/10.14214/sf.513
Keywords: mixed models; berry yield index; Vaccinium myrtillus; Vaccinium vitis-idaea
Abstract | View details | Full text in PDF | Author Info
Forest berries and the outdoor experiences related to berry collection are important goods and services provided by Finnish forests. Consequently, there is a need for models which facilitate the prediction of the impacts of alternative forest management options on berry yields. Very few such models are available. In particular, empirical models are lacking. Models used in forest management should express the effect of variables altered in forest management such as stand density and mean tree size. This study developed empirical models for bilberry and cowberry yields in North Karelia. The data consisted of 362 measurements of 40 m2 sample plots. The plots were located in clusters. The same plot was measured over 1 to 4 years. Besides berry yield some site and growing stock characteristics of each plot were measured. A random parameter model was used to express the berry yield as a function of site fertility, growing stock characteristics, and random parameters. The random part of the models accounted for the effect of plot, measurement year, and cluster. The fixed predictors of the model for bilberry were stand age and forest site type. Stand basal area, mean tree diameter and forest site type were used to predict cowberry yields. The most significant random parameter was the plot factor. The fixed model part explained only a few per cent of the variation in berry yields. The signs of regression coefficients were logical and the model predictions correlated rather well with the predictions of earlier models.
  • Ihalainen, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: marjut.ihalainen@joensuu.fi (email)
  • Salo, The Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: ks@nn.fi
  • Pukkala, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 524, category Research article
Timo Pukkala, Jari Miina, Marc Palahí. (2002). Thinning response and thinning bias in a young Scots pine stand. Silva Fennica vol. 36 no. 4 article id 524. https://doi.org/10.14214/sf.524
Keywords: Pinus sylvestris; growth model; non-spatial model; spatial model
Abstract | View details | Full text in PDF | Author Info
The study analyses the annual post-thinning response and thinning bias of a young Scots pine stand as a function of tree size, competition faced by the tree, and competition that is removed around the tree in the thinning treatment. The thinning response of a tree was defined as the change of tree growth due to a thinning treatment. The thinning bias was defined as the difference between the true growth and model prediction. A distance-dependent (spatial) and a distance-independent (non-spatial) growth model were used in the calculations. The empirical data were measured from a thinning experiment consisting of ten plots, each 40 x 30 m in size, which were thinned to different stand densities. The ten-year post-thinning growth of every remaining tree was measured. The results indicated that the highest thinning response is among medium-sized and co-dominant trees. The thinning response is quite small, and even negative for some trees, for two years after thinning but it becomes clearly positive from the third year onwards. The spatial model underestimated the growth of small trees (which usually face high competition) while the non-spatial model overestimated the growth of trees that are small or face much competition. The spatial model used in this study overemphasized the effect of competition while the non-spatial model underestimated this effect. Both growth models overestimated the growth of trees in heavily thinned places, but this bias disappeared in two years. The negative bias was more pronounced with a spatial growth model because the tendency of the non-spatial model to underestimate the growth of trees facing little competition partly compensated for the negative bias.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: jm@nn.fi
  • Palahí, European Forest Institute, Torikatu 34, FIN-80100 Joensuu, Finland E-mail: mp@nn.fi
article id 588, category Research article
Marjut Ihalainen, Timo Pukkala. (2001). Modelling cowberry (Vaccinium vitis-idaea) and bilberry (Vaccinium myrtillus) yields from mineral soils and peatlands on the basis of visual field estimates. Silva Fennica vol. 35 no. 3 article id 588. https://doi.org/10.14214/sf.588
Keywords: Vaccinium myrtillus; Vaccinium vitis-idaea; non-wood forest products
Abstract | View details | Full text in PDF | Author Info
This study presents new models for predicting bilberry and cowberry yields from site and stand characteristics. These models enable one to evaluate the future states of forests in terms of berry yields. The modelling data consisted of visual field estimates of site and tree stand characteristics, as well as berry yields from 627 forest stands. Berry yields were estimated using a scale from 0 to 10. Using these data, models were prepared which predict the berry yield scores from those site and stand characteristics which are usually known in forest planning calculations. The model predictions correlated positively and often quite strongly with earlier models. The results were in line with previous studies on the effects of site and tree cover on berry production. According to the models, sites of medium and rather poor fertility produce the highest bilberry yields. Increasing tree height increases, and the basal area of spruce and proportion of deciduous trees decrease, bilberry yield. With mineral soils, cowberry yields are best on poor sites. A high proportion of pine improves cowberry yields. The yields are the highest in open areas and very young stands, on the one hand, and in sparsely populated stands of large and old trees, on the other hand. In pine swamps, the yields are best on rather poor sites. Increasing basal area of deciduous trees decreases cowberry yields.
  • Ihalainen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: marjut.ihalainen@forest.joensuu.fi (email)
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 621, category Research article
Jyrki Kangas, Pekka Leskinen, Timo Pukkala. (2000). Integrating timber price scenario modeling with tactical management planning of private forestry at forest holding level. Silva Fennica vol. 34 no. 4 article id 621. https://doi.org/10.14214/sf.621
Keywords: decision support; optimization; adaptive behaviour; non-industrial private forestry; tactical planning; timber price modelling
Abstract | View details | Full text in PDF | Author Info
In forest management planning, deterministic timber prices are typically assumed. However, real-life timber prices vary in the course of time, and also price peaks, i.e. exceptionally high timber prices, might occur. If land-owners can utilise the price variation by selling timber with the high prices, they are able to increase their net revenues correspondingly. In this study, an approach is presented to study the timber price variation and its significance in the optimization of forest management. The approach utilizes stochastic timber price scenario modelling, simulation of forest development, and optimization of forest management. The approach is presented and illustrated by means of a case study. It is shown how the degree of uncertainty due to variation in timber prices can be analyzed in tactical forest planning of private forestry, and how the potential benefits of adaptive timber-selling behaviour for a forest landowner can be computed by using the approach. The effects of stochastic timber prices on the choice of forest plan are studied at the forest holding level considering also the spacing and type of cuttings and the optimal cutting order. A forest plan prepared under the assumption of constant timber price very seldom results in optimal forest management. Through studying the effects of stochastic timber prices, forest landowners and other decision makers obtain valuable information about the significance of adaptive timber selling behaviour. The presented methodology can also be used in analysing the land-owners’ economic risks as a function of time-price structure.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jyrki.kangas@metla.fi (email)
  • Leskinen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: pl@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi

Category : Research note

article id 310, category Research note
Timo Tahvanainen, Kalle Kaartinen, Timo Pukkala, Matti Maltamo. (2007). Comparison of approaches to integrate energy wood estimation into the Finnish compartment inventory system. Silva Fennica vol. 41 no. 1 article id 310. https://doi.org/10.14214/sf.310
Keywords: diameter distribution; thinnings; calibration estimation; compartment inventory
Abstract | View details | Full text in PDF | Author Info
The harvesting of energy wood from young stands is increasing as the demand for renewable wood fuel is growing. Energy wood consists of stems, tree tops, branches and needles, depending on the size of the trees and the logging method used. The current forest inventory and planning systems used in private forests in Finland do not produce estimates of energy wood components. In stands typical for energy wood harvesting, a large share of energy wood consists of trees smaller than the minimum size for pulpwood. In this study, energy wood was included into the calculation system of compartment inventory, and a procedure for simulating the thinning treatments in young stands was developed. The results for six inventory alternatives and prediction of energy wood were compared with the use of inventory material from 37 young stands that have plenty of energy wood. The measurement of additional stand characteristics and the use of a calibration estimation method was tested, as well as the use of plot-level inventory data instead of stand level data. The results showed that the measurement of the number of trees per hectare, in addition to stand basal area and mean diameter, improved the energy wood estimates. The additional minimum and maximum diameters improved the precision of the estimates, but did not affect bias. The removal estimates were more precise when plot-level data was used, rather than stand-level data. The removal estimates were higher with plot-level data. The results suggest that, in heterogeneous young stands, plot by plot prediction would give more accurate removal estimates than the calculation of a corresponding prediction at the stand-level.
  • Tahvanainen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: timo.tahvanainen@metla.fi (email)
  • Kaartinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: kk@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: tp@nn.fi
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: mm@nn.fi

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